Echis Carinatus Multisquamatus
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''Echis carinatus multisquamatus'', known as the multiscale saw-scaled viper and transcaspian saw-scaled viper,
a
Wildlife of Pakistan
Accessed 3 August 2006.
is a
viper The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs tha ...
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
found in
Uzbekistan Uzbekistan (, ; uz, Ozbekiston, italic=yes / , ; russian: Узбекистан), officially the Republic of Uzbekistan ( uz, Ozbekiston Respublikasi, italic=yes / ; russian: Республика Узбекистан), is a doubly landlocked cou ...
,
Turkmenistan Turkmenistan ( or ; tk, Türkmenistan / Түркменистан, ) is a country located in Central Asia, bordered by Kazakhstan to the northwest, Uzbekistan to the north, east and northeast, Afghanistan to the southeast, Iran to the sout ...
,
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
,
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan,; prs, امارت اسلامی افغانستان is a landlocked country located at the crossroads of Central Asia and South Asia. Referred to as the Heart of Asia, it is bordere ...
, and
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
. Like all other vipers, it is
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
.


Description

This subspecies grows to in total length (body + tail), but usually no more than .Mallow D, Ludwig D, Nilson G. 2003. ''True Vipers: Natural History and Toxinology of Old World Vipers''. Malabar, Florida: Krieger Publishing Company. 359 pp. . Its head marking is always cross-shaped. Its lateral white line is continuous and undulating. Narrow transverse white bands occur on the middorsum. The
dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...
are in 34-40 rows at midbody, the highest number of rows of all subspecies of ''E. carinatus''. The
ventrals In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...
number 169-199 (highest of all subspecies).


Geographic range

The range for this snake is from
Uzbekistan Uzbekistan (, ; uz, Ozbekiston, italic=yes / , ; russian: Узбекистан), officially the Republic of Uzbekistan ( uz, Ozbekiston Respublikasi, italic=yes / ; russian: Республика Узбекистан), is a doubly landlocked cou ...
, south to
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
, and east to western
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
. Th
Wildlife of Pakistan
website describes the range as from the
Caspian Sea The Caspian Sea is the world's largest inland body of water, often described as the world's largest lake or a full-fledged sea. An endorheic basin, it lies between Europe and Asia; east of the Caucasus, west of the broad steppe of Central Asia ...
, through
Turkmenistan Turkmenistan ( or ; tk, Türkmenistan / Түркменистан, ) is a country located in Central Asia, bordered by Kazakhstan to the northwest, Uzbekistan to the north, east and northeast, Afghanistan to the southeast, Iran to the sout ...
and Uzbekistan (
Bukhara Bukhara (Uzbek language, Uzbek: /, ; tg, Бухоро, ) is the List of cities in Uzbekistan, seventh-largest city in Uzbekistan, with a population of 280,187 , and the capital of Bukhara Region. People have inhabited the region around Bukhara ...
,
Samarkand fa, سمرقند , native_name_lang = , settlement_type = City , image_skyline = , image_caption = Clockwise from the top:Registan square, Shah-i-Zinda necropolis, Bibi-Khanym Mosque, view inside Shah-i-Zinda, ...
,
Tashkent Tashkent (, uz, Toshkent, Тошкент/, ) (from russian: Ташкент), or Toshkent (; ), also historically known as Chach is the capital and largest city of Uzbekistan. It is the most populous city in Central Asia, with a population of ...
), into
Tajikistan Tajikistan (, ; tg, Тоҷикистон, Tojikiston; russian: Таджикистан, Tadzhikistan), officially the Republic of Tajikistan ( tg, Ҷумҳурии Тоҷикистон, Jumhurii Tojikiston), is a landlocked country in Centr ...
, along the
Afghan Afghan may refer to: *Something of or related to Afghanistan, a country in Southern-Central Asia *Afghans, people or citizens of Afghanistan, typically of any ethnicity ** Afghan (ethnonym), the historic term applied strictly to people of the Pas ...
border up to
Hindukush The Hindu Kush is an mountain range in Central and South Asia to the west of the Himalayas. It stretches from central and western Afghanistan, Quote: "The Hindu Kush mountains run along the Afghan border with the North-West Frontier Provinc ...
, northeastern
Balochistan Balochistan ( ; bal, بلۏچستان; also romanised as Baluchistan and Baluchestan) is a historical region in Western and South Asia, located in the Iranian plateau's far southeast and bordering the Indian Plate and the Arabian Sea coastline. ...
,
Khyber-Pakhtunkhwa Khyber Pakhtunkhwa (; ps, خېبر پښتونخوا; Urdu, Hindko: خیبر پختونخوا) commonly abbreviated as KP or KPK, is one of the four provinces of Pakistan. Located in the northwestern region of the country, Khyber Pakhtunkhwa ...
, and eastern Iran. The type locality is listed as "Bairam-Ali, Mariskaya oblast, Turkmeniya" (
Baýramaly Baýramaly (formerly Bayram-Ali, tk, Baýramaly, earlier Bahrām Ali ) is a city in and the seat of Baýramaly District, Mary Province, Turkmenistan. It lies about 27 km east of the provincial capital Mary, along the main railway line from ...
,
Mary Region Mary Region ( tk, Mary welaýaty, Мары велаяты) is one of five provinces in Turkmenistan. It is located in the south-east of the country, bordering Afghanistan. Its capital is the city of Mary. Its area is and population 1,480,400 ( ...
,
Turkmenistan Turkmenistan ( or ; tk, Türkmenistan / Түркменистан, ) is a country located in Central Asia, bordered by Kazakhstan to the northwest, Uzbekistan to the north, east and northeast, Afghanistan to the southeast, Iran to the sout ...
).


Taxonomy

It was classified as a separate
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
, ''E. multisquamatus'', when first described by Cherlin (1981).


References


Further reading

* Auffenberg W, Rehman H. 1991. Studies on Pakistan Reptiles. Pt. 1. The genus ''Echis'' (Viperidae). ''Bull. Florida Mus. Nat. Hist., Biol Sci.'' 35 (5): 263-314. * Cherlin VA. 1981. new saw-scaled viper, ''Echis multisquamatus sp. nov.'', from southwestern and central Asia 'Proc. Zool. Inst. Acad. Sci. USSR''101: 92-95. (in Russian).


External links

* * {{Taxonbar, from=Q5332603 Viperinae Reptiles of Pakistan