Echeveria Amphoralis
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''Echeveria'' is a large
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Crassulaceae The Crassulaceae (from Latin ''crassus'', thick), also known as the stonecrop family or the orpine family, are a diverse family of dicotyledon flowering plants characterized by succulent leaves and a unique form of photosynthesis, known as Crass ...
,
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to semi-desert areas of Central America, Mexico and northwestern South America.


Description

Plants may be
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
or
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
. Flowers on short stalks (cymes) arise from compact rosettes of
succulent In botany, succulent plants, also known as succulents, are plants with parts that are thickened, fleshy, and engorged, usually to retain water in arid climates or soil conditions. The word ''succulent'' comes from the Latin word ''sucus'', meani ...
fleshy, often brightly coloured leaves. Species are
polycarpic Polycarpic plants are those that flower and set seeds many times before dying. A term of identical meaning is pleonanthic and iteroparous. Polycarpic plants are able to reproduce multiple times due to at least some portion of its meristems being ab ...
, meaning that they may flower and set seed many times over the course of their lifetimes. Often numerous
offset Offset or Off-Set may refer to: Arts, entertainment, and media * "Off-Set", a song by T.I. and Young Thug from the '' Furious 7: Original Motion Picture Soundtrack'' * ''Offset'' (EP), a 2018 EP by singer Kim Chung-ha * ''Offset'' (film), a 200 ...
s are produced, and are commonly known as "
hen and chicks Hen and chicks (also known as hen-and-chickens, or hen-widdies in the southern United States) is a common name for a group of small succulent plants, a term that indicates a plant that possesses enlarged parts to store water. It belongs to th ...
", which can also refer to other genera, such as ''
Sempervivum ''Sempervivum'' (Brit. , U.S. ©sɛ̃mpeÉč'vivĆ©m is a genus of about 40 species of flowering plants in the family Crassulaceae, commonly known as houseleeks. Other common names include liveforever (the source of the taxonomical designation ''Sem ...
'', that are significantly different from ''Echeveria''. Many species of ''Echeveria'' serve important environmental roles, such as those of
host plant In biology and medicine, a host is a larger organism that harbours a smaller organism; whether a parasitic, a mutualistic, or a commensalist ''guest'' ( symbiont). The guest is typically provided with nourishment and shelter. Examples include ...
s for butterflies. For example, the butterfly ''Callophrys xami'' uses several species of ''Echeveria'', such as ''Echevelia gibbiflora'', for suitable host plants. Even more, these plants are integral to the oviposition process of ''C. xami'' and some other butterfly species as well.


Etymology

''Echeveria'' is named for
Atanasio EcheverrĂ­a y Godoy Atanasio EcheverrĂ­a y Godoy was an 18th-century Mexican people, Mexican botanical artist and naturalist who trained at the Royal Art Academy in Mexico. The genus ''Echeveria'' was named in his honour by Augustin Pyramus de Candolle. Royal Botani ...
, a botanical illustrator who contributed to ''Flora Mexicana''.Gledhill, David (2008). "The Names of Plants". Cambridge University Press. (hardback), (paperback). p. 149


Taxonomy

The genus was erected by
A. P. de Candolle Augustin Pyramus (or Pyrame) de Candolle (, , ; 4 February 17789 September 1841) was a Swiss botanist. René Louiche Desfontaines launched de Candolle's botanical career by recommending him at a herbarium. Within a couple of years de Candolle ...
in 1828, and is named after the 18th century Mexican botanical artist
Atanasio EcheverrĂ­a y Godoy Atanasio EcheverrĂ­a y Godoy was an 18th-century Mexican people, Mexican botanical artist and naturalist who trained at the Royal Art Academy in Mexico. The genus ''Echeveria'' was named in his honour by Augustin Pyramus de Candolle. Royal Botani ...
. , the genus consists of about 150 species, including genera such as ''Oliveranthus'' and ''Urbinia'' that have formerly been split off from ''Echeveria''.
Molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
studies have repeatedly shown the genus not to be
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
: species of ''Echeveria'' cluster with species of '' Cremnophila'', ''
Graptopetalum ''Graptopetalum'' (leatherpetal) is a plant genus of the family ''Crassulaceae''. They are perennial succulent plants and native to Mexico and Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, AlÄ­ áčŁonak ) is a state in the Southwestern ...
'', ''
Pachyphytum Pachyphytum is a small genus of succulents in the family Crassulaceae, native to Mexico Mexico (Spanish: MĂ©xico), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north b ...
'', and '' Thompsonella'' as well as species of ''
Sedum ''Sedum'' is a large genus of flowering plants in the family Crassulaceae, members of which are commonly known as stonecrops. The genus has been described as containing up to 600 species, subsequently reduced to 400–500. They are leaf succulen ...
'' sect. ''Pachysedum''. The former ''Urbinia'' species do appear to form a monophyletic group within this grouping. Although it is clear that ''Echeveria'' is not monophyletic, its limits are not clear, and further analyses are needed to determine whether and how the genus should be split, or if it should be included in an expanded concept of ''
Sedum ''Sedum'' is a large genus of flowering plants in the family Crassulaceae, members of which are commonly known as stonecrops. The genus has been described as containing up to 600 species, subsequently reduced to 400–500. They are leaf succulen ...
''.


Species

One source accepts the following species:


Cultivation

Many ''Echeveria'' species are popular as
ornamental garden plants Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that i ...
. They are drought-resistant, although they do better with regular deep watering and fertilizing. Most will tolerate shade and some frost, although
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
s tend to be less tolerant. Most lose their lower leaves in winter; as a result, after a few years, the plants lose their compact appearance and need to be re-rooted or propagated. In addition, if not removed, the shed leaves may decay, harboring fungus that can then infect the plant.


Propagation

They can be propagated easily by separating offsets, but also by leaf cuttings, and by
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
if they are not hybrids.


Cultivars and Hybrids

''Echeveria'' has been extensively bred and hybridized. The following is a selection of available plants. *‘Arlie Wright’ *‘Black Prince’ *‘Blue Heron’ *‘Blue Surprise’ (''E.'' × ''gilva'') *'Crimson Tide' *‘Dondo’ *‘Doris Taylor’ *‘Ebony’ (''E. agavoides'' cultivar) *‘Frank Reinelt’ *‘Hoveyi’ *‘Lipstick’ (''E. agavoides'' cultivar) *‘Oliver’ (''E. pulvinata'' cultivar) *‘Opalina’ *‘Painted Lady’ *‘Perle von NĂŒrnberg’ *‘Paul Bunyan’ *‘Red’ (''E.'' × ''gilva'') *‘Ruberia’ *‘Set-Oliver’ *‘Tippy’ *‘Victor Reiter’ (''E. agavoides'' cultivar) *‘Wavy Curls’ *‘Worfield Wonder’ (''E.'' × ''derosa'')


Award winning

The following have gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nort ...
’s
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
: *''Echeveria agavoides'' *''Echeveria chihuahuaensis'' *''Echeveria derenbergii'' *''Echeveria elegans'' *''Echeveria'' ‘Perle von NĂŒrnberg’ *''Echeveria runyonii'' ‘Topsy Turvy’ *''Echeveria secunda'' var. ''glauca'' ‘Compton Carousel’ *''Echeveria setosa'' *''Echeveria'' × ''bombycina''


Formerly in ''Echeveria''

*''
Dudleya ''Dudleya'', commonly known as liveforevers (Spanish: ''siemprevivas'') is a genus of succulent plants in the stonecrop family, Crassulaceae, consisting of about 68 taxa in southwestern North America and Guadalupe Island. The species come in mul ...
'' –
Alwin Berger Alwin Berger (28 August 1871 – 20 April 1931) was a German botanist best known for his contribution to the nomenclature of succulent plants, particularly agaves and cacti. Born in Germany he worked at the botanical gardens in Dresden and Fran ...
placed this as a subtaxon of ''Echeveria'', and species within were described as ''Echeveria''. *'' Graptopetalum paraguayense'' (N.E.Br.) E.Walther (as ''E. weinbergii'' hort. ex T.B.Sheph.) *'' Pachyveria clavifolia'' (as ''E. clavifolia'') *'' Reidmorania occidentalis'' (as ''E. kimnachii'')


Photo gallery

File:Echeveria Blue Curl.jpg, Flowers of ''Echeveria'' ‘Blue Curl’ File:Echeveria 'Black Prince' - 20090226.jpg, ''Echeveria'' cv. ‘Black Prince’ File:Echeveria pulvinata smith.jpg, The flowers of '' E. pulvinata'' are in bright yellowish orange color. (MHNT) Echeveria secunda.jpg, '' Echeveria secunda'' File: é»‘çŽ‹ć­çš„èŠ±ćș20190417202338.jpg , Echeveria ‘Black Prince’ . 2018 Taichung World Flora Exposition, Taiwan. File:Succulent vertical wall.jpg, Sculptural quality of Echeveria on display File:Echeveria Compton Carousel.jpg, Echeveria "Compton Carousel"


References


External links


''Flora Mexicana'' via Biodiversity Library
{{Taxonbar, from=Q837077 Crassulaceae genera Succulent plants Taxa named by Augustin Pyramus de Candolle