Eastern Wood Pewee
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The eastern wood pewee (''Contopus virens'') is a small
tyrant flycatcher The tyrant flycatchers (Tyrannidae) are a family of passerine birds which occur throughout North and South America. They are considered the largest family of birds known to exist in the world, with more than 400 species. They are the most dive ...
from
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. This
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
and the
western wood pewee The western wood pewee (''Contopus sordidulus'') is a small tyrant flycatcher. Adults are gray-olive on the upperparts with light underparts, washed with olive on the breast. They have two wing bars and a dark bill with yellow at the base of the ...
(''C. sordidulus'') were formerly considered a single
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
. The two species are virtually identical in appearance, and can be distinguished most easily by their calls.


Taxonomy

In 1760 the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published works ...
included a description of the eastern wood pewee in his ''Ornithologie'' based on a specimen collected in the
Carolinas The Carolinas are the U.S. states of North Carolina and South Carolina, considered collectively. They are bordered by Virginia to the north, Tennessee to the west, and Georgia to the southwest. The Atlantic Ocean is to the east. Combining Nort ...
. He used the French name ''Le gobe-mouche cendré de la Coroline'' and the Latin ''Muscicapa Carolinensis cinerea''. Although Brisson coined Latin names, these do not conform to the binomial system and are not recognised by the
International Commission on Zoological Nomenclature The International Commission on Zoological Nomenclature (ICZN) is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 26 commissioners from 20 countries. Orga ...
. When in 1766 the Swedish naturalist
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
updated his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' for the twelfth edition, he added 240 species that had been previously described by Brisson. One of these was the eastern wood pewee. Linnaeus included a brief description, coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Muscicapa virens'' and cited Brisson's work. The specific name ''virens'' is Latin for "green". This species is now placed in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Contopus'' that was introduced by the German ornithologist
Jean Cabanis Jean Louis Cabanis (8 March 1816 – 20 February 1906) was a German ornithologist. Cabanis was born in Berlin to an old Huguenot family who had moved from France. Little is known of his early life. He studied at the University of Berlin from 18 ...
in 1855. The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
.


Description

The eastern wood pewee is in length and weighs around . Wingspan ranges from 9.1-10.2 in (23-26 cm). The sexes are alike. The adult is gray-olive on the upperparts with light underparts, washed with olive on the breast. Each wing has two pale wing bars, and the primary
remiges Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...
are long, giving the wingtip a slim and very pointed appearance. The upper part of the bill is dark, the lower part is yellowish. The songs are basically a mournful whistled ''pee-a'wee'' given in a series, which gave this bird its name, and a "pe-wee" with a rising note at the end. The
western wood pewee The western wood pewee (''Contopus sordidulus'') is a small tyrant flycatcher. Adults are gray-olive on the upperparts with light underparts, washed with olive on the breast. They have two wing bars and a dark bill with yellow at the base of the ...
(''C. sordidulus'') is essentially indistinguishable visually. But its range is
parapatric In parapatric speciation, two subpopulations of a species evolve reproductive isolation from one another while continuing to exchange genes. This mode of speciation has three distinguishing characteristics: 1) mating occurs non-randomly, 2) gene ...
to the west of the eastern wood pewee and its song—a descending ''tsee-tsee-tsee-peeer''—is entirely different. The
eastern phoebe The eastern phoebe (''Sayornis phoebe'') is a small passerine bird. The genus name ''Sayornis'' is constructed from the specific part of Charles Lucien Bonaparte's name for Say's phoebe, ''Muscicapa saya'', and Ancient Greek ''ornis'', "bird". ...
(''Sayornis phoebe'') is similar, particularly in the worn plumage after breeding. It always lacks clearly defined wingbars, however, and bobs its tail frequently. It has a shorter primary projection. The eastern phoebe is also present on the breeding grounds by March, while eastern wood pewees do not arrive until very late April and early May. The songs (''fee-bee, fee-bee'') and calls (''chip'') are quite different. The
least flycatcher The least flycatcher (''Empidonax minimus'') (also called chebec, or chebecker, after the sound it makes) is a small insect-eating bird. It is the smallest ''Empidonax'' flycatcher in eastern North America. Taxonomy The closest relative to the ...
(''Empidonax minimus'') is quite similar to the eastern wood pewee in plumage, but has a bold eye ring and much shorter primary projection, appearing rather blunt-winged. It also has a shorter bill and is smaller overall. The songs (''che-bec, che-bec'') and calls (a sharp ''whit'') are very different.


Distribution and habitat

Their breeding
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
is
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
, mixed woods, or pine plantations in eastern
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. They migrate to
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
, the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
, and in the
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
region of northern
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
. They feed on
insects Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of j ...
and other
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s. Wood pewees wait on a perch at a middle height in a tree and fly out to catch prey in flight, sometimes hovering to pick it from vegetation. Eastern wood pewees arrive relatively late on breeding grounds (e.g., 18 May to 5 June in southern Ontario). They are rarely seen on their breeding grounds before the last days of April further south. They migrate south at a more usual time, leaving sometimes in late August but most often in September. Migration times have stayed the same in the last 100 years. They migrate quickly for most of the journey, dispersing and moving at a slower speed when approaching breeding or wintering range. For example, in the
Cayman Islands The Cayman Islands () is a self-governing British Overseas Territory—the largest by population in the western Caribbean Sea. The territory comprises the three islands of Grand Cayman, Cayman Brac and Little Cayman, which are located to the ...
which are offshore the winter quarters, the species is only rarely and briefly encountered, but passes through on its way north as early as mid-late February.


Behaviour


Breeding

The eastern wood pewee makes an open
cup nest A bird nest is the spot in which a bird lays and incubates its eggs and raises its young. Although the term popularly refers to a specific structure made by the bird itself—such as the grassy cup nest of the American robin or Eurasian bl ...
made of
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...
es, bark, and
lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...
s (''Quercus''),
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...
s (''Pinus''),
birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech-oak family Fagaceae. The genus ''Betula'' contains 30 ...
es (''Betula''), and
maple ''Acer'' () is a genus of trees and shrubs commonly known as maples. The genus is placed in the family Sapindaceae.Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 9, June 2008 nd more or less continuously updated since http ...
s (''Acer''). The female lays almost always three (sometimes two) translucent-white eggs with brown flecking concentrated towards the larger end of the ovate egg. Males are territorial and defend the nesting area aggressively, often fighting with neighbouring conspecifics and even pursue attacks on other species (e.g., least flycatchers, American robins, chipping sparrows, red-eyed vireos, etc.). Males can sometimes be
polygynous Polygyny (; from Neoclassical Greek πολυγυνία (); ) is the most common and accepted form of polygamy around the world, entailing the marriage of a man with several women. Incidence Polygyny is more widespread in Africa than in any ...
, mating with two females, simultaneously. The eggs hatch in 12–14 days and both parents bring food to the
altricial In biology, altricial species are those in which the young are underdeveloped at the time of birth, but with the aid of their parents mature after birth. Precocial species are those in which the young are relatively mature and mobile from the mome ...
nestlings. Nestlings typically fledge 15–17 days after hatching, often ending up on the ground during the first flight out of the nest. The adults will perch on a nearby branch and call out to the nestlings, keeping contact and providing them with food until the young are able to fly to join them.


Conservation status

The eastern wood pewee is common and widespread, and therefore not considered globally threatened by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
. Its numbers, however, are declining in recent decades, possibly due to the loss of forest habitat in its winter range. It is also possible that the increase of
white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known as the whitetail or Virginia deer, is a medium-sized deer native to North America, Central America, and South America as far south as Peru and Bolivia. It has also been introduced t ...
(''Odocoileus virginianus'') in its breeding range has led to a change in vegetation and associated
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s in the lower levels of the deciduous forests where the eastern wood pewee breeds. File:Contopus virensPCCA20050724-9873B.jpg, Eastern wood pewee, has two crisp, pale wing bars and long primary projection File:WesternWoodPewee23.jpg,
Western wood pewee The western wood pewee (''Contopus sordidulus'') is a small tyrant flycatcher. Adults are gray-olive on the upperparts with light underparts, washed with olive on the breast. They have two wing bars and a dark bill with yellow at the base of the ...
looks like its
sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...
File:Sayornis phoebe CT4.jpg,
Eastern phoebe The eastern phoebe (''Sayornis phoebe'') is a small passerine bird. The genus name ''Sayornis'' is constructed from the specific part of Charles Lucien Bonaparte's name for Say's phoebe, ''Muscicapa saya'', and Ancient Greek ''ornis'', "bird". ...
lacks bolder wing bars and has shorter primary projection File:Empidonax-minimus-001.jpg,
Least flycatcher The least flycatcher (''Empidonax minimus'') (also called chebec, or chebecker, after the sound it makes) is a small insect-eating bird. It is the smallest ''Empidonax'' flycatcher in eastern North America. Taxonomy The closest relative to the ...
has bold eye-rings and shorter primary projection


References


External links


Eastern wood pewee – ''Contopus virens''
– USGS Patuxent Bird Identification InfoCenter

– Cornell Lab of Ornithology
Bird sound
- Florida Museum of Natural History

Familiar Birds

- Canadian Biodiversity website
Eastern wood pewee
Environment Canada * * * {{Authority control
eastern wood pewee The eastern wood pewee (''Contopus virens'') is a small tyrant flycatcher from North America. This bird and the western wood pewee (''C. sordidulus'') were formerly considered a single species. The two species are virtually identical in appeara ...
Native birds of the Eastern United States Birds of Central America Birds of the Caribbean
eastern wood pewee The eastern wood pewee (''Contopus virens'') is a small tyrant flycatcher from North America. This bird and the western wood pewee (''C. sordidulus'') were formerly considered a single species. The two species are virtually identical in appeara ...
eastern wood pewee The eastern wood pewee (''Contopus virens'') is a small tyrant flycatcher from North America. This bird and the western wood pewee (''C. sordidulus'') were formerly considered a single species. The two species are virtually identical in appeara ...