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The Dartford warbler (''Curruca undata'') is a typical warbler from the warmer parts of western
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
and northwestern
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
. It is a small warbler with a long thin tail and a thin pointed bill. The adult male has grey-brown upperparts and is dull reddish-brown below except for the centre of the belly which has a dirty white patch. It has light speckles on the throat and a red eye-ring. The sexes are similar but the adult female is usually less grey above and paler below. Its breeding range lies west of a line from southern
England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separa ...
to the heel of
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
(southern
Apulia it, Pugliese , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = , demographics1_footnotes = , demographics1_title1 = , demographics1_info1 = , demographic ...
). The Dartford warbler is usually resident all year in its breeding range, but there is some limited
migration Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum l ...
.


Taxonomy and systematics

The Dartford warbler was first described in 1776 by the Welsh naturalist,
Thomas Pennant Thomas Pennant (14 June OS 172616 December 1798) was a Welsh naturalist, traveller, writer and antiquarian. He was born and lived his whole life at his family estate, Downing Hall near Whitford, Flintshire, in Wales. As a naturalist he h ...
. He introduced the English name and based his description on two specimens that had been obtained by the ornithologist John Latham from Bexley Heath, near
Dartford Dartford is the principal town in the Borough of Dartford, Kent, England. It is located south-east of Central London and is situated adjacent to the London Borough of Bexley to its west. To its north, across the Thames estuary, is Thurrock in ...
in
Kent Kent is a county in South East England and one of the home counties. It borders Greater London to the north-west, Surrey to the west and East Sussex to the south-west, and Essex to the north across the estuary of the River Thames; it fac ...
. In 1783 Latham included the warbler in his ''A General Synopsis of Birds'' but did not coin the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
, ''Sylvia dartfordiensis'', until the supplement to his work was published in 1787. However, in 1783, before the publication of Latham's supplement, the Dutch naturalist
Pieter Boddaert Pieter Boddaert (1730 – 6 May 1795) was a Dutch physician and naturalist. Early life, family and education Boddaert was the son of a Middelburg jurist and poet by the same name (1694–1760). The younger Pieter obtained his M.D. at the Univers ...
introduced the name, ''Motacilla undata'', based on a coloured plate of "Le Pitte-chou, de Provence" in Edmé-Louis Daubenton's ''Planches enluminées d'histoire naturelle''. The specific epithet ''undata'' is from
Medieval Latin Medieval Latin was the form of Literary Latin used in Roman Catholic Western Europe during the Middle Ages. In this region it served as the primary written language, though local languages were also written to varying degrees. Latin function ...
''undatus'' meaning "with wavy markings". The type locality is
Provence Provence (, , , , ; oc, Provença or ''Prouvènço'' , ) is a geographical region and historical province of southeastern France, which extends from the left bank of the lower Rhône to the west to the Italian border to the east; it is bor ...
in France. This species probably forms a superspecies with
Tristram's warbler Tristram's warbler (''Curruca deserticola'') is a species of Old World warbler in the family Sylviidae. It is found in Algeria, Libya, Mauritania, Morocco, Tunisia, and Western Sahara. Its natural habitat is subtropical dry shrubland. The spe ...
and this in turn seems close to Marmora's warbler and the
Balearic warbler Balearic warbler (''Curruca balearica'') is a typical warbler, genus ''Curruca''. It is endemic to the Balearic Islands, apart from Menorca. It groups with the Marmora's warbler, Tristram's warbler and the Dartford warbler (Helbig 2001, Jønss ...
. Altogether, this group of typical warblers bears a resemblance to the
wrentit The wrentit (''Chamaea fasciata'') is a small bird that lives in chaparral, oak woodlands, and bushland on the western coast of North America. It is the only species in the genus ''Chamaea''. Its systematics have been the subject of much debat ...
, the only species of
Sylviidae Sylviidae is a family of passerine birds that includes the typical warblers and a number of babblers formerly placed within the Old World babbler family. They are found in Eurasia and Africa. Taxonomy and systematics The scientific name Syl ...
from the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with t ...
. However, the wrentit is less closely related to the genus ''Sylvia'' than to the parrotbills. Its visual similarity to the Dartford warbler group is an example of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
. Three
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised: * ''C. u. dartfordiensis'' ( Latham, 1787) – south England and north-west France * ''C. u. toni'' ( Hartert, 1909) – north-west Africa * ''C. u. undata'' ( Boddaert, 1783) –
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
and south France


Description

The Dartford warbler is a small, ,
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
bird, distinguished by its long tail compared with that of other warblers. Its plumage comprises unobtrusive and muted tones, which blend in with the dry dead plants, old wood or sunny greyish wood found in its preferred habitats. Like many typical warblers, the Dartford warbler has distinct male and female
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
s. The male has a grey back and head, reddish underparts, and a red eye. The reddish throat is spotted with white. The sides are a dull greyish tone, being more clear about the abdomen. In some populations males have bluish-grey or brownish-grey backs and heads. The female is paler below, especially on the throat, and a browner grey above. The female's throat also has white spots, although they are smaller and less marked than in the male. Juvenile birds are similar to females.


Distribution and habitat

The species is naturally rare. The largest European populations of ''Curruca undata'' are in the Iberian peninsula, others in much of
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
, in
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
and southern
England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separa ...
and south
Wales Wales ( cy, Cymru ) is a country that is part of the United Kingdom. It is bordered by England to the east, the Irish Sea to the north and west, the Celtic Sea to the south west and the Bristol Channel to the south. It had a population in ...
. In
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
it can be found only in small areas in the north, wintering in northern
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
and northern
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...
.


Behaviour and ecology


Breeding

Dartford warblers first breed when they are one year old. They are usually monogamous and the
pair bond In biology, a pair bond is the strong affinity that develops in some species between a mating pair, often leading to the production and rearing of offspring and potentially a lifelong bond. Pair-bonding is a term coined in the 1940s that is freque ...
can persist from one year to the next. The male sometimes builds several simple nests (cock nests) of which one is chosen by the female, but it is more usual for both birds to participate in constructing the breeding nest. In southern England the birds breed on heathlands, sometimes near the coast, and nest in either
common gorse ''Ulex europaeus'', the gorse, common gorse, furze or whin, is a species of flowering plant in the family Fabaceae, native to the British Isles and Western Europe. Description Growing to tall, it is an evergreen shrub. The young stems are ...
(''Ulex europaeus'') or common heather (''Calluna vulgaris''). The compact cup-shaped nest is located in dense bushes, usually within of the ground. It is formed mainly of grasses and is lined with a layer of finer material that can include thin roots and feathers. The eggs are laid from early April in southern France and Spain, and from mid-April in southern England. The
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts) ...
is typically 3–5 eggs which are smooth and glossy, with a white or occasionally pale green ground and marked with brown speckles which are sometimes concentrated at the larger end. The average size of an egg is with a weight of of which 6 percent is shell. The eggs are incubated for 12–14 days mainly by the female. The chicks are fed
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, ...
s by both parents. The nestlings
fledge Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable ...
10–14 days after hatching and are then fed by their parents for a further two weeks. Usually two and occasionally three broods are raised in a year.


Feeding

It inhabits open fields with degraded scrub brush and is common in heather. In winter it may visit urban areas, but always feeds within shrubs in these areas. It nests in bushes with thorns and near the ground. These warblers are mostly
insectivorous A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores we ...
, eating
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder S ...
s,
butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises ...
,
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 descri ...
s and
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species div ...
s. The song of the Dartford warbler is a distinctive rattling warble.


Status and conservation

Dartford warblers almost died out in the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and No ...
in the severe winter of 1962/1963 when the national population dropped to just ten pairs. ''Sylvia undata'' is also sensitive to drought affecting breeding success or producing heath fires, as occurred during 1975 and 1976 in England when virtually all juveniles failed to survive their first year.Bibby, Colin J.
‘Dartford Warblers in England’
''British Birds'', 72 (1979); pp. 10–22
However, this species can recover well in good quality habitat with favourable temperatures and rainfall, thanks to repeated nesting and a high survival rate for the young. Indeed, they recovered in some areas of the UK, but numbers are once again on the decline in other regions of their natural range. The range of the Dartford warbler is restricted to western and southern Europe. The total population in 2012 was estimated at 1.1–2.5 million breeding pairs. The largest numbers occur in Spain where there were believed to be 983,000–1,750,000 pairs. For reasons that probably include loss of suitable habitat, the Spanish population appears to be declining. The species is therefore classed by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of nat ...
as being
Near Threatened A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify f ...
. A period of climatic warming since 1963 has seen the UK population increase to "more than 2,500 pairs in 2006 (Wotton et al. 2009). Expansion into patches of structurally suitable habitat (up to an altitude of 400m), more northerly areas and away from the core of the range, from Dorset and Hampshire to Derbyshire and Suffolk, is likely to have been facilitated by milder winter weather (Wotton et al. 2009, Bradbury et al. 2011). The Dartford warbler population in the UK is expected to continue to increase. However, future climate-based projections for the European range indicate that by 2080, more than 60% of the current European range may no longer be suitable (Huntley et al 2007). There is evidence that this is happening already, with severe declines in Spain and France (Green 2017). For this reason, the species is classified as Near Threatened on the IUCN Global Red List. If the declines in southern Europe continue, the UK will become increasingly important for global conservation of this species".


References


Sources

*
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
(BLI) (2008)
2008 IUCN Redlist status changes
Retrieved 23 May 2008 *


External links


Ageing and sexing (PDF; 3.1 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze

Photos and videos from the Internet Bird Collection
* *
Vocalisations on Xeno-canto
{{Taxonbar, from=Q110257524, from2=Q752545
Dartford warbler The Dartford warbler (''Curruca undata'') is a typical warbler from the warmer parts of western Europe and northwestern Africa. It is a small warbler with a long thin tail and a thin pointed bill. The adult male has grey-brown upperparts and is d ...
Birds of Europe Birds of North Africa
Dartford warbler The Dartford warbler (''Curruca undata'') is a typical warbler from the warmer parts of western Europe and northwestern Africa. It is a small warbler with a long thin tail and a thin pointed bill. The adult male has grey-brown upperparts and is d ...