Dytiscus Avunculus
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''Dytiscus'' ("little diver" based on Greek ''δυτικός'', "able to dive" and the diminutive suffix ''-ίσκος'') is a
Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical region ...
genus of
predaceous diving beetle The Dytiscidae – based on the Greek ''dytikos'' (δυτικός), "able to dive" – are the predaceous diving beetles, a Family (biology), family of water beetles. They occur in virtually any freshwater habitat around the world, but a ...
s that usually live in
wetland A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...
s and
pond A pond is an area filled with water, either natural or artificial, that is smaller than a lake. Defining them to be less than in area, less than deep, and with less than 30% emergent vegetation helps in distinguishing their ecology from th ...
s. There are 26 species in this genus distributed in Europe, Asia, North Africa and North and Central America. They are predators that can reduce mosquito
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e. ''Dytiscus'' are large water beetles with a robust, rounded shape and they measure long depending on the exact species involved. The largest, '' D. latissimus'', is among the largest species in the family and its size is only matched by certain ''
Megadytes ''Megadytes'' is a genus of diving beetles in the family Dytiscidae. They are found in slow-moving or static freshwater habitats throughout most of the Neotropics, ranging from Florida and Mexico, through the West Indies and Central America, to S ...
''. The tarsi of the males are modified into suckers which are used to grip the female in mating. Females are usually larger than the males and come in two forms, with grooved (sulcate) or smooth elytra. Males only ever have smooth elytra. The adults of most species can fly.


Life history

Adult beetles and their larvae are aquatic but the pupae spend their life in the ground. Females lay eggs inside the tissue of aquatic plants such as reeds. The eggs hatch in about three weeks. The larvae (known as "water tigers") are elongate with a round and flat head and strong mandibles. They are predatory and their mandible have grooves on their inner edge through which they are able to suck the body fluids of their prey. The larvae take air from the surface of the water using hairs at the end of their abdomen. These lead to
spiracle Spiracle or spiraculum may refer to: * Spiracle (arthropods), opening in the exoskeletons of some arthropods * Spiracle (vertebrates), openings on the surface of some vertebrates * Spiraculum, a genus of land snails in family Cyclophoridae Cycl ...
s into which the air is taken. Once the larvae grow to some size, they move to soil at the edge of water and burrow into a cell and pupate. The adults breathe by going to the surface and upending. They collect air under their elytra and are able to breathe this collected air using spiracles hidden under the elytra. In '' Dytiscus marginalis'' and other species the tarsus of the forelegs is modified in males to form a circular sucker. A reduced sucker is also seen in the midleg of the male.


Parasitoids

Eggs of ''Dytiscus'' are sometimes parasitized by wasps of the families
Eulophidae The Eulophidae are a large family of hymenopteran insects, with over 4,300 described species in some 300 genera. The family includes the genus ''Elasmus'', which used to be treated as a separate family, "Elasmidae", and is now treated as a subf ...
,
Mymaridae The Mymaridae, commonly known as fairyflies or fairy wasps, are a family (biology), family of chalcid wasp, chalcidoid wasps found in temperate and tropical regions throughout the world. The family contains around 100 genera with 1400 species. F ...
and other
Chalcidoidea Chalcid wasps (, , for their metallic colour) are insects within the superfamily Chalcidoidea, part of the order Hymenoptera. The superfamily contains some 22,500 known species, and an estimated total diversity of more than 500,000 species, me ...
.


Species

''Dysticus'' contains the following species:Dytiscidae Species List
at Joel Hallan's Biology Catalog. Texas A&M University. Retrieved on 7 May 2012.
* '' Dytiscus alaskanus'' J.Balfour-Browne, 1944 * '' Dytiscus avunculus'' C.Heyden, 1862 * '' Dytiscus caraboides'' Linnaeus,
1758 Events January–March * January 1 – Swedish biologist Carl Linnaeus (Carl von Linné) publishes in Stockholm the first volume (''Animalia'') of the 10th edition of Systema Naturae, 10th edition of ''Systema Naturae'', the sta ...
* '' Dytiscus carolinus'' Aubé, 1838 * '' Dytiscus circumcinctus'' (Ahrens, 1811) * ''
Dytiscus circumflexus ''Dytiscus'' ("little diver" based on Greek ''δυτικός'', "able to dive" and the diminutive suffix ''-ίσκος'') is a Holarctic genus of predaceous diving beetles that usually live in wetlands and ponds. There are 26 species in this gen ...
'' Fabricius, 1801 * '' Dytiscus cordieri'' Aubé, 1838 * ''
Dytiscus dauricus ''Dytiscus dauricus'' is a species of predaceous diving beetle in the family Dytiscidae The Dytiscidae – based on the Greek ''dytikos'' (δυτικός), "able to dive" – are the predaceous diving beetles, a family of water beetle ...
'' Gebler, 1832 * ''
Dytiscus delictus ''Dytiscus'' ("little diver" based on Greek ''δυτικός'', "able to dive" and the diminutive suffix ''-ίσκος'') is a Holarctic genus of predaceous diving beetles that usually live in wetlands and ponds. There are 26 species in this genu ...
'' (Zaitzev, 1906) * ''
Dytiscus dimidiatus ''Dytiscus'' ("little diver" based on Greek ''δυτικός'', "able to dive" and the diminutive suffix ''-ίσκος'') is a Holarctic genus of predaceous diving beetles that usually live in wetlands and ponds. There are 26 species in this genu ...
'' Bergsträsser, 1778 * ''
Dytiscus distantus ''Dytiscus'' ("little diver" based on Greek ''δυτικός'', "able to dive" and the diminutive suffix ''-ίσκος'') is a Holarctic genus of predaceous diving beetles that usually live in wetlands and ponds. There are 26 species in this genu ...
'' Feng, 1936 * ''
Dytiscus fasciventris ''Dytiscus fasciventris'', the understriped diving beetle, is a species of predaceous diving beetle in the family Dytiscidae The Dytiscidae – based on the Greek ''dytikos'' (δυτικός), "able to dive" – are the predaceous divi ...
'' Say, 1824 * ''
Dytiscus habilis ''Dytiscus habilis'' is a species of predaceous diving beetle in the family Dytiscidae. It is found in North America from Oklahoma to Arizona south throughout Mexico to Guatemala; it is the only ''Dytiscus'' species found in the Neotropics T ...
'' Say, 1830 * '' Dytiscus harrisii'' Kirby, 1837 * '' Dytiscus hatchi'' Wallis, 1950 * '' Dytiscus hybridus'' Aubé, 1838 * '' Dytiscus krausei'' H.J.Kolbe, 1931 * '' Dytiscus lapponicus'' Gyllenhal, 1808 * '' Dytiscus latahensis'' Wickham, 1931 * ''
Dytiscus latissimus ''Dytiscus latissimus'' is a species of beetle in family Dytiscidae. Description One of the largest representatives of the predaceous diving beetles of the genus '' Dytiscus'', ''D. latissimus'' can reach a length around . This beetle is similar ...
'' Linnaeus,
1758 Events January–March * January 1 – Swedish biologist Carl Linnaeus (Carl von Linné) publishes in Stockholm the first volume (''Animalia'') of the 10th edition of Systema Naturae, 10th edition of ''Systema Naturae'', the sta ...
* '' Dytiscus latro'' Sharp, 1882 * '' Dytiscus lavateri'' Heer, 1847 * '' Dytiscus marginalis'' Linnaeus,
1758 Events January–March * January 1 – Swedish biologist Carl Linnaeus (Carl von Linné) publishes in Stockholm the first volume (''Animalia'') of the 10th edition of Systema Naturae, 10th edition of ''Systema Naturae'', the sta ...
* '' Dytiscus marginicollis'' LeConte, 1845 * '' Dytiscus miocenicus'' Lewis & Gundersen, 1987 * '' Dytiscus mutinensis'' Branden, 1885 * '' Dytiscus persicus'' Wehncke, 1876 * '' Dytiscus pisanus'' Laporte, 1835 * '' Dytiscus semisulcatus'' (O.F.Müller, 1776) * '' Dytiscus sharpi'' Wehncke, 1875 * '' Dytiscus sinensis'' Feng, 1935 * '' Dytiscus thianschanicus'' (Gschwendtner, 1923) * '' Dytiscus verticalis'' Say, 1823 * '' Dytiscus zersii'' Sordelli, 1882 Image:DytiscusEgg.jpg, Eggs and egglaying Image:DytiscusLarva.jpg, Larva Image:DytiscusPupa.png, Pupa Image:DytiscusSpiracles.png, Spiracles under the elytra Image:DytiscusMale.png, Male with suckers on fore tarsi Image:DytiscusLifeLyd.png, Life history


References


External links

* {{Taxonbar, from=Q1138850 Dytiscidae genera