Dunlin

The dunlin (''Calidris alpina'') is a small wader, formerly sometimes separated with the other "stints" in the genus ''Erolia''. The English name is a dialect form of "dunling", first recorded in 1531–1532. It derives from ''dun'', "dull brown", with the suffix ''-ling'', meaning a person or thing with the given quality.

It is a circumpolar breeder in Arctic or subarctic regions. Birds that breed in northern Europe and Asia are long-distance migrants, wintering south to Africa, southeast Asia and the Middle East. Birds that breed in Alaska and the Canadian Arctic migrate short distances to the Pacific and Atlantic coasts of North America, although those nesting in northern Alaska overwinter in Asia. Many dunlins winter along the Iberian south coast.

Taxonomy

The dunlin was formally described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his ''Systema Naturae'' under the binomial name ''Tringa alpina''. Linnaeus specified the location as Lapland. This species was formerly placed in the genus ''Erolia'', but is now placed with 23 other sandpipers in the genus ''Calidris'' that was introduced in 1804 by the German naturalist Blasius Merrem. The genus name is from Ancient Greek ''kalidris'' or ''skalidris'', a term used by Aristotle for some grey-coloured waterside birds. The specific ''alpina '' is from Latin and means "of high mountains", in this case referring to the Alps.

Ten subspecies are recognised:
*''C. a. arctica'' (Schiøler, 1922) — breeds in northeast Greenland
*''C. a. schinzii'' ( Brehm & Schilling, 1822) — breeds in southeast Greenland, Iceland, the British Isles, Scandinavia & the Baltic
*''C. a. alpina'' (Linnaeus, 1758) — breeds in northern Europe and northwest Siberia
*''C. a. centralis'' (Buturlin, 1932) — breeds in north-central and northeast Siberia
*''C. a. sakhalina'' (Vieillot, 1816) — breeds in eastern Russia to the Chukchi Peninsula
*''C. a. kistchinski'' Tomkovich, 1986 — breeds around the Sea of Okhotsk to Kuril Islands and Kamchatka
*''C. a. actites'' Nechaev & Tomkovich, 1988 — breeds on Sakhalin
*''C. a. arcticola'' (Todd, 1953) — breeds from northwest Alaska to northwest Canada
*''C. a. pacifica'' (Coues, 1861) — breeds in western and southern Alaska
*''C. a. hudsonia'' (Todd, 1953) — breeds in central Canada

Description

Measurements:

* Length:
* Weight:
* Wingspan:

An adult dunlin in breeding plumage shows the distinctive black belly which no other similar-sized wader possesses. The winter dunlin is basically grey above and white below. Juveniles are brown above with two whitish "V" shapes on the back. They usually have black marks on the flanks or belly and show a strong white wingbar in flight. The legs and slightly decurved bill are black. There are a number of subspecies differing mainly in the extent of rufous colouration in the breeding plumage and the bill length. Bill length varies between sexes, the females having longer bills than the males. On the tip of the Dunlin's bill is a soft covering that fills with blood and with many nerve endings, forming a sensitive probe that is used to locate invertebrate prey in mud and sand. Although the bill can look sharp-pointed in dead specimens, in life it is blunt.

The call is a typical sandpiper "peep", and the display song a harsh trill.

Distribution and habitat

Dunlin are small migratory waders, however they show strong philopatry with individuals of the Southern Dunlin (''Calidris alpina schinzii'') in Sweden and Finland returning to, or very close to, their natal patches. Habitat fragmentation has reduced the availability of habitat patches to these birds through reducing patch size and increasing patch isolation. This reduced connectivity between patches has reduced the movements of Dunlin leaving them more susceptible to inbreeding in these locations. Future management for the conservation of Southern Dunlin should include increasing the connectivity between habitat patches.

Behaviour

The dunlin is highly gregarious in winter, sometimes forming large flocks on coastal mudflats or sandy beaches. Large numbers can often be seen swirling in synchronized flight on stop-overs during migration or on their winter habitat.

This bird is one of the most common waders throughout its breeding and wintering ranges, and it is the species with which other waders tend to be compared. At length and with a wingspan, it is similar in size to a common starling, but stouter, with a thicker bill.

The dunlin moves along the coastal mudflat beaches it prefers with a characteristic "sewing machine" feeding action, methodically picking small food items. Insects form the main part of the dunlin's diet on the nesting grounds; it eats molluscs, worms and crustaceans in coastal areas.

Breeding

The nest is a shallow scrape on the ground lined with vegetation, into which typically four eggs are laid and incubated by the male and female parents. Chicks are precocial, however are brooded during early development. They start to fly at approximately three weeks of age. The majority of brood care is provided by the male, as the female deserts the brood and often leaves the breeding area.

Apparent hybrids between this species and the white-rumped sandpiper as well as with the purple sandpiper have been reported from the Atlantic coasts of North America and Europe, respectively.

Status

The dunlin has an extremely large range and although the population appears to be decreasing, the population is still very large. The International Union for Conservation of Nature (IUCN) has judged that the threat to the species is of "Least concern". The dunlin is one of the species to which the ''Agreement on the Conservation of African-Eurasian Migratory Waterbirds'' (AEWA) applies.

Gallery

File:Seasonal-Variations-in-the-Diet-and-Foraging-Behaviour-of-Dunlins-Calidris-alpina-in-a-South-pone.0081174.s001.ogv, Foraging
File:Kärrsnäppa - (Calidris alpina).jpg, Ystad, Sweden
File:Alpenstrandläufer (calidris alpina) - Spiekeroog, Nationalpark niedersächsisches Wattenmeer.jpg, Adult in breeding plumage, Spiekeroog, northern Germany
File:Calidris alpina pibydd y mawn.jpg, Adult in winter plumage, Wales
File:Calidris alpina alpina, Riga, Latvia 1.jpg, Adult in breeding plumage, Latvia

References

External links

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Dunlin Species Account
- Cornell Lab of Ornithology


- USGS Patuxent Bird Identification InfoCenter

Ageing and sexing (PDF; 1.6 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze
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Calidris
Birds of the Arctic
Holarctic birds
Birds of the Dominican Republic
Birds of the Caribbean
Birds of North Africa
Birds described in 1758
Taxa named by Carl Linnaeus
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