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''Dryopteris affinis'', the scaly male fern or golden-scaled male fern, is a
fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to western and southern Europe and southwestern Asia. It is most abundant on moist soils in woodlands in areas with high humidity, such as the British Isles and western France. In the Mediterranean region and the Caucasus it is confined to high altitudes.


Description

''Dryopteris affinis'' is virtually
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
and bears light green
frond A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the lar ...
s long, moderately stiff and hard-textured, the
rachis In biology, a rachis (from the grc, ῥάχις [], "backbone, spine") is a main axis or "shaft". In zoology and microbiology In vertebrates, ''rachis'' can refer to the series of articulated vertebrae, which encase the spinal cord. In this c ...
at the base of the frond densely covered in yellow-brown scales known as ramenta. The frond is bi
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
, the pinnae up to long, the pinnules broad rectangular with the margin most toothed close to the pinna tip. There is a blackish spot at the base of the pinna where it joins the rachis. Individual fronds live for about 1.5 years and remain attached to the
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
after withering. ''D. affinis'' is closely related to ''
Dryopteris filix-mas ''Dryopteris filix-mas'', the male fern, is a common fern of the temperate Northern Hemisphere, native to much of Europe, Asia, and North America. It favours damp shaded areas in the understory of woodlands, but also shady places on hedge-banks, ...
'', distinguished by its usually more robust habit with usually more evergreen fronds, more densely scaly frond stems, and more rectangular (less tapered and lobed) pinnae and pinnules. It is one of the larger European native ferns, with older specimens developing a dense, almost
tree fern The tree ferns are arborescent (tree-like) ferns that grow with a trunk elevating the fronds above ground level, making them trees. Many extant tree ferns are members of the order Cyatheales, to which belong the families Cyatheaceae (scaly tree ...
-like base up to high and broad.


Cultivation

Numerous
cultivars A cultivar is a type of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and when Plant propagation, propagated retain those traits. Methods used to propagate cultivars include: division, root and st ...
and
varieties Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
have been selected for garden use, of which the following have gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nort ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
: *''D. affinis'' *'Polydactyla Mapplebeck' *'Crispa Gracilis' *'Cristata' *'Cristata Angustata' Dryopteris affinis1.jpg, Young fronds in spring Ramenta on Dryopteris affinis.JPG, Ramenta on the fern's rachis Dryopteris affinis - 'shuttlecock'.JPG, 'Shuttlecock' form of growth


References


External links


Flora Europaea: ''Dryopteris affinis''
*Hyde, H. A., Wade, A. E., & Harrison, S. G. (1978). ''Welsh Ferns''. National Museum of Wales. {{Taxonbar, from=Q2036293 affinis Ferns of Asia Ferns of Europe Flora of Iran Flora of Western Asia Flora of England Flora of Norway Taxa named by Richard Thomas Lowe