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''Banksia wonganensis'' is a large
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
endemic to
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
that, until 2007, was previously known as ''Dryandra wonganensis''. It occurs within a small area in the vicinity of the
Wongan Hills Wongan Hills is a range of low flat-topped hills in the Avon Wheatbelt bioregion of Western Australia. It is located at , in the Shire of Wongan–Ballidu. History The range was first recorded in 1836 by Surveyor General of Western Australia Jo ...
. It grows on
lateritic Laterite is both a soil and a rock type rich in iron and aluminium and is commonly considered to have formed in hot and wet tropical areas. Nearly all laterites are of rusty-red coloration, because of high iron oxide content. They develop by ...
soils in open woodland or amongst dense shrub. It is rare, but does not appear to be endangered.


Description

It is a sprawling or erect shrub, up to high, without a
lignotuber A lignotuber is a woody swelling of the root crown possessed by some plants as a protection against destruction of the plant stem, such as by fire. Other woody plants may develop basal burls as a similar survival strategy, often as a response t ...
. Leaves are long, and nine to 18 millimetres wide, and
pinnatifid Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common Anatomical terms of location#Axes, axis. Pinnation occurs in biological morphology (biology), morphology, in Crysta ...
, with four to nine triangular lobes on each side, before tapering to a fine point. They are often strongly curved. Flowers are bright yellow, and occur in heads of 45 to 50 flowers, up to across. Each flower consists of a tubular
perianth The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower, and structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepals when ...
made up of four fused
tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s, and one long wiry
style Style is a manner of doing or presenting things and may refer to: * Architectural style, the features that make a building or structure historically identifiable * Design, the process of creating something * Fashion, a prevailing mode of clothing ...
; the head of the style is initially trapped inside the upper perianth parts, but breaks free at
anthesis Anthesis is the period during which a flower is fully open and functional. It may also refer to the onset of that period. The onset of anthesis is spectacular in some species. In ''Banksia'' species, for example, anthesis involves the extension ...
. After flowering, follicles develop in the woody base of the flower head, each containing one or sometimes two seeds.


Taxonomy

Early collections of ''B. wonganensis'' include specimens collected by Alexander Morrison at Wongan Hills in October 1903, by Norm Moyle in Monk Well Gully, Wongan Hills, by Fred Lullfitz northwest of Wongan Hills, and by Kenneth Newbey in the Wongan Hills. In 1985 these specimens were wrongly attributed to ''Dryandra hewardiana'' (now '' Banksia hewardiana'') by Robert Malcolm Sainsbury in his ''Field Guide to Dryandras'', but it was later recognised as a distinct species, and referred to in
FloraBase ''FloraBase'' is a public access web-based database of the flora of Western Australia. It provides authoritative scientific information on 12,978 taxa, including descriptions, maps, images, conservation status and nomenclatural details. 1,272 alie ...
as ''Dryandra'' sp. 25. The type specimen was collected by Alex George on 4 August 1986, from a location on Piawaning Road north-west of Wongan Hills. George published a formal description of the species in 1996 in ''
Nuytsia ''Nuytsia floribunda'' is a hemiparasitic tree found in Western Australia. The species is known locally as moodjar and, more recently, the Christmas tree or Western Australian Christmas tree. The display of intensely bright flowers during the ...
'' 10(3). He placed it in the genus ''
Dryandra ''Banksia'' ser. ''Dryandra'' is a series of 94 species of shrub to small tree in the plant genus ''Banksia''. It was considered a separate genus named ''Dryandra'' until early 2007, when it was merged into ''Banksia'' on the basis of extensiv ...
'', subgenus ''Dryandra'', series ''Armatae'', and gave it the specific epithet "wonganensis" from "wongan", after the Wongan Hills where it is found, and the Latin "ensis" (''origin'' or ''place''). Thus its full name was ''Dryandra wonganensis'' A.S.George. Early in 2007,
Austin Mast Austin R. Mast is a research botanist. Born in 1972, he obtained a Ph.D. from the University of Wisconsin–Madison in 2000. He is currently a professor within the Department of Biological Science at Florida State University (FSU), and has been dire ...
and
Kevin Thiele Kevin R. Thiele is currently an adjunct associate professor at the University of Western Australia and the director of Taxonomy Australia. He was the curator of the Western Australian Herbarium from 2006 to 2015. His research interests include ...
transferred all ''Dryandra'' taxa to ''Banksia''. The current name for this species is therefore ''Banksia wonganensis'' (A.S.George) A.R.Mast & K.R.Thiele. As an interim measure, Mast and Thiele placed all but one ''Dryandra'' taxon in ''Banksia'' ser. ''Dryandra''.


Distribution and habitat

''Banksia wonganensis'' occurs only in the
Wongan Hills Wongan Hills is a range of low flat-topped hills in the Avon Wheatbelt bioregion of Western Australia. It is located at , in the Shire of Wongan–Ballidu. History The range was first recorded in 1836 by Surveyor General of Western Australia Jo ...
and on surrounding rises, in the
Avon Wheatbelt The Avon Wheatbelt is a bioregion in Western Australia. It has an area of . It is considered part of the larger Southwest Australia savanna ecoregion. Geography The Avon Wheatbelt bioregion is mostly a gently undulating landscape with low reli ...
biogeographic Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
region. The location has
lateritic Laterite is both a soil and a rock type rich in iron and aluminium and is commonly considered to have formed in hot and wet tropical areas. Nearly all laterites are of rusty-red coloration, because of high iron oxide content. They develop by ...
soils covered by open woodland or dense scrub. Mean annual rainfall is , with a mean temperature range of , and up to 80 days above .


Ecology

Like most other
Proteaceae The Proteaceae form a family of flowering plants predominantly distributed in the Southern Hemisphere. The family comprises 83 genera with about 1,660 known species. Together with the Platanaceae and Nelumbonaceae, they make up the order Pro ...
, ''B. wonganensis'' has
proteoid root Cluster roots, also known as proteoid roots, are plant roots that form clusters of closely spaced short lateral rootlets. They may form a two- to five-centimetre-thick mat just beneath the leaf litter. They enhance nutrient uptake, possibly by chem ...
s, roots with dense clusters of short lateral rootlets that form a mat in the soil just below the leaf litter. These enhance solubilisation of nutrients, thus allowing nutrient uptake in low-nutrient soils such as the
phosphorus Phosphorus is a chemical element with the symbol P and atomic number 15. Elemental phosphorus exists in two major forms, white phosphorus and red phosphorus, but because it is highly reactive, phosphorus is never found as a free element on Ear ...
-deficient native soils of Australia. It lacks a lignotuber, so is killed by bushfire. However it is adapted to release its seed following a fire, so populations regenerate rapidly. It has been given a rating of "Priority Four - Poorly Known Taxa" on Western Australia's Department of Environment and Conservation's
Declared Rare and Priority Flora List The Declared Rare and Priority Flora List is the system by which Western Australia's conservation flora are given a priority. Developed by the Government of Western Australia's Department of Environment and Conservation, it was used extensively wi ...
, meaning that it has been adequately surveyed, has been determined to be rare, but is apparently not threatened.


Cultivation

The species is little known in cultivation. Tony Cavanagh and
Margaret Pieroni Margaret Pieroni is a Western Australian botanical artist Botanical illustration is the art of depicting the form, color, and details of plant species, frequently in watercolor paintings. They must be scientifically accurate but often also ha ...
rate its foliage as attractive and its flowers as showy, and state that it is just as attractive as many other large shrubs in its series. The species prefer heavy soils with good drainage, and tolerates full sun or light shade. It has good tolerance for both drought and frost. Germination of seed appears to have a success rate of only about 50%; germination is rapid, often taking less than four weeks.


References


External links

* * * {{Taxonbar, from=Q4856765 wonganensis Endemic flora of Western Australia Taxa named by Kevin Thiele