Dromopus
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''Dromopus'' is a reptilian ichnogenus commonly found in assemblages of ichnofossils dating to the late
Pennsylvanian Pennsylvanian may refer to: * A person or thing from Pennsylvania * Pennsylvanian (geology) The Pennsylvanian ( , also known as Upper Carboniferous or Late Carboniferous) is, in the International Commission on Stratigraphy, ICS geologic timesca ...
(
Moscovian Moscovian may refer to: *An inhabitant of Moscow, the capital of Russia *Something of, from, or related to Moscow *Moscovian (Carboniferous) The Moscovian is in the International Commission on Stratigraphy, ICS geologic timescale a stage (strati ...
stage) to the late
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...
(
Changhsingian In the geologic time scale, the Changhsingian or Changxingian is the latest age or uppermost stage of the Permian. It is also the upper or latest of two subdivisions of the Lopingian Epoch or Series. The Changhsingian lasted from to 251.902 mill ...
stage). It has been found throughout Europe, as well as in the United States, Canada, and Morocco. Several ichnospecies have been named; only the type ichnospecies ''D. lacertoides'' is definitively recognized.


History


Species

Originally named as an ichnospecies of '' Saurichnites'' by
Hanns Bruno Geinitz Hanns Bruno Geinitz (16 October 1814 – 28 January 1900) was a German geologist, born at Altenburg, the capital of Saxe-Altenburg. He was educated at the universities of Berlin and Jena, and gained the foundations of his geological knowledge ...
in 1861, ''S. lacertoides'' was transferred to the newly created genus ''Dromopus'' by
Othniel Charles Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of Paleontology in Yale College and President of the National Academy of Sciences. He was one of the preeminent scientists in the field of paleontology. Among h ...
in 1894, along with a new ichnospecies ''D. agilis'' from the United States. In 1929, Roy Moodie described two-digit traces from the Red Beds of Texas under the ichnospecies '' Varanopus palmatus'', ''V. impressus'', ''V. elrodi'', and ''V. didactylus''. However, these were subsequently considered to represent a single ichnospecies of ''Dromopus'' by
William Sarjeant William Antony Swithin Sarjeant (15 July 1935 – 8 July 2002), also known by the pen name Antony Swithin, was a professor of geology at University of Saskatchewan. He was also interested in mystery writing, fantasy writing, folk singing, and h ...
. Because the
type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...
of ''V. palmatus'' is broken, and ''V. impressus'' and ''V. elrodi'' are incomplete, ''Dromopus didactylus'' is generally the preferred name. In 1963 and 1964, Paul Ellenberger described many ichnospecies of ''Dromopus'': ''D. neooctavianensis'', ''D. brevidigitatus'', ''D. duidigitatus'', ''D. octavianensis'', ''D. bidigitatus'', ''D. curtidigitus'', ''D. inversidigitulifer'', ''D. rabejacensis'', ''D. rectidigitus'', ''D. rectangulus'', ''D. exsultans'', and ''D. cursor''. Harmut Haubold considered them invalid in a 2000 review. There have been disagreements about whether the North American ''D. agilis'' represents a separate ichnospecies. More recently, based on the fact that there is no satisfactory anatomical diagnosis distinguishing various ichnospecies of ''Dromopus'', only the type ichnospecies ''D. lacertoides'' has been agreed to be valid.


Synonyms

In 1892, Pohlig used the new ichnogenus ''Protritonichnites'' for the ichnospecies ''Saurichnites lacertoides'', before Marsh had named ''Dromopus''. By the principles of synonymy, ''Protritonichnites'' should have priority over ''Dromopus''. However, in 1970, Haubold argued that this name was largely unused by this point, and it was also based on a less taxonomically sound definition. Although some European researchers during the 1980s persisted in using the name ''Protritonichnites'', ''Dromopus'' has generally found more widespread adoption. Other synonyms include ''Eumekichnium'', named by
Franz Nopcsa Franz may refer to: People * Franz (given name) * Franz (surname) Places * Franz (crater), a lunar crater * Franz, Ontario, a railway junction and unorganized town in Canada * Franz Lake, in the state of Washington, United States – see ...
in 1923, including the type ichnospecies ''E. longipollex'', as well as ''E. gampsodactylum'' and ''E. pachydactylum'' (both formerly ''Ichnium''); and ''Gampsodactylum'', also named by Nopcsa in 1923, including the type ichnospecies ''G. alberdorfense'' as well as ''G. friedrichrodanum'' and ''G. kabarzense''.


Description

''Dromopus lacertoides'' tracks are characterized by five-fingered hand and foot imprints up to long, about a third longer than they are wide, with a short palm and long, slender, tapered digits. The length of the digits increases from the first to the fourth, with the fifth having the same length as the second. The hand and foot imprints are not particularly distinct. ''D. didactylus'' tracks are based on similar tracks, but where the imprints of all but two digits are faint. ''Dromopus'' tracks have been ascribed to lizard-like reptiles, including the diapsid Araeoscelidia and the parareptilian Bolosauridae.


References

{{Taxonbar, from=Q96376648 Reptile trace fossils