Dillenia Pentagyna
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A small tree with tortuous twigs, ''Dillenia pentagyna'' is a member of the family
Dilleniaceae Dilleniaceae is a family of flowering plants with 11 genera and about 430 known species. Such a family has been universally recognized by taxonomists. It is known to gardeners for the genus ''Hibbertia'', which contains many commercially valuabl ...
, and is found from Sulawesi to South-Central China to India and Sri Lanka. Material from the tree has some minor uses.


Description

A tree some 6-15m tall, with tortuous twigs, the bark is grayish and smooth, exfoliating. Branches are glabrous and stout. Leaves are deciduous, petiolate, oblong to obovate-oblong, glabrous, 30–5 cm long, flowers appear before the leaves, 2–7 in number, yellow coloured petals, flowering starts in April–May. Fruit is globose, 0.5 cm in diameter, black ovoid seed, exarillate. On islands of the
Mekong The Mekong or Mekong River is a trans-boundary river in East Asia and Southeast Asia. It is the world's List of rivers by length, twelfth longest river and List of longest rivers of Asia, the third longest in Asia. Its estimated length is , ...
in northeastern
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand t ...
, the tree flowers from February to March, fruits from March to April, and the leaves grow from May to November.


Vernacular names

Common names by which the tree is known include: ''buku''
Tetun , nativename=Tetun , states= Indonesia East Timor , speakers=, mostly in Indonesia , date=2010–2011 , ref=e18 , speakers2=50,000 L2-speakers in Indonesia and East Timor , familycolor=Austronesian , fam2=Malayo-Polynesian , fam3= Central–East ...
,
Timor Timor is an island at the southern end of Maritime Southeast Asia, in the north of the Timor Sea. The island is East Timor–Indonesia border, divided between the sovereign states of East Timor on the eastern part and Indonesia on the western p ...
; ''janti'', ''sempu'' Jawa; ''l've/lve'', ''chhë muëy rô:y'', ''pheng'', ''rovey'', ''muoy roy sratoap'' Khmer; 小花五桠果;, ''xiao hua wu ya guo''
Chinese Chinese can refer to: * Something related to China * Chinese people, people of Chinese nationality, citizenship, and/or ethnicity **''Zhonghua minzu'', the supra-ethnic concept of the Chinese nation ** List of ethnic groups in China, people of va ...
; ส้านช้าง
Thai Thai or THAI may refer to: * Of or from Thailand, a country in Southeast Asia ** Thai people, the dominant ethnic group of Thailand ** Thai language, a Tai-Kadai language spoken mainly in and around Thailand *** Thai script *** Thai (Unicode block ...
; ''korkotta''
Bengali Bengali or Bengalee, or Bengalese may refer to: *something of, from, or related to Bengal, a large region in South Asia * Bengalis, an ethnic and linguistic group of the region * Bengali language, the language they speak ** Bengali alphabet, the w ...
; ''akshi''
Assam Assam (; ) is a state in northeastern India, south of the eastern Himalayas along the Brahmaputra and Barak River valleys. Assam covers an area of . The state is bordered by Bhutan and Arunachal Pradesh to the north; Nagaland and Manipur ...
; ''aggai'', ''kallai''
Hindi Hindi (Devanāgarī: or , ), or more precisely Modern Standard Hindi (Devanagari: ), is an Indo-Aryan language spoken chiefly in the Hindi Belt region encompassing parts of northern, central, eastern, and western India. Hindi has been de ...
; ''aksikiphal'', ''punnaga''
Sanskrit Sanskrit (; attributively , ; nominally , , ) is a classical language belonging to the Indo-Aryan branch of the Indo-European languages. It arose in South Asia after its predecessor languages had diffused there from the northwest in the late ...
; ''agaai'', ''taatarii'', ''tatri chinnakalinga'' Nepali; ''karmal''
Gujarati Gujarati may refer to: * something of, from, or related to Gujarat, a state of India * Gujarati people, the major ethnic group of Gujarat * Gujarati language, the Indo-Aryan language spoken by them * Gujarati languages, the Western Indo-Aryan sub- ...
and
Marathi Marathi may refer to: *Marathi people, an Indo-Aryan ethnolinguistic group of Maharashtra, India *Marathi language, the Indo-Aryan language spoken by the Marathi people *Palaiosouda, also known as Marathi, a small island in Greece See also * * ...
; ''rai'' Oriya; ''kanigala'', ''kadu-kanigala''
Kannada Kannada (; ಕನ್ನಡ, ), originally romanised Canarese, is a Dravidian language spoken predominantly by the people of Karnataka in southwestern India, with minorities in all neighbouring states. It has around 47 million native s ...
; പട്ടിപ്പുന്ന], ''punna'', ''kodapunna''
Malayalam Malayalam (; , ) is a Dravidian language spoken in the Indian state of Kerala and the union territories of Lakshadweep and Puducherry (Mahé district) by the Malayali people. It is one of 22 scheduled languages of India. Malayalam was des ...
; ''ravudana''
Telugu Telugu may refer to: * Telugu language, a major Dravidian language of India *Telugu people, an ethno-linguistic group of India * Telugu script, used to write the Telugu language ** Telugu (Unicode block), a block of Telugu characters in Unicode S ...
; பஞ்சகன்னி உவா, ''naytekku'', ''kanigala''
Tamil Tamil may refer to: * Tamils, an ethnic group native to India and some other parts of Asia ** Sri Lankan Tamils, Tamil people native to Sri Lanka also called ilankai tamils **Tamil Malaysians, Tamil people native to Malaysia * Tamil language, nati ...
; ''dog teak''
English English usually refers to: * English language * English people English may also refer to: Peoples, culture, and language * ''English'', an adjective for something of, from, or related to England ** English national ide ...
; ''dillenia''; ''Nepali elephant apple''


Distribution

The tree grows in the following places:
Lesser Sunda Islands The Lesser Sunda Islands or nowadays known as Nusa Tenggara Islands ( id, Kepulauan Nusa Tenggara, formerly ) are an archipelago in Maritime Southeast Asia, north of Australia. Together with the Greater Sunda Islands to the west they make up t ...
including
Timor Timor is an island at the southern end of Maritime Southeast Asia, in the north of the Timor Sea. The island is East Timor–Indonesia border, divided between the sovereign states of East Timor on the eastern part and Indonesia on the western p ...
,
Sulawesi Sulawesi (), also known as Celebes (), is an island in Indonesia. One of the four Greater Sunda Islands, and the world's eleventh-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Mindanao and the Sulu Ar ...
, Jawa,
Peninsular Malaysia Peninsular Malaysia ( ms, Semenanjung Malaysia; Jawi: سمننجڠ مليسيا), or the States of Malaya ( ms, Negeri-negeri Tanah Melayu; Jawi: نڬري-نڬري تانه ملايو), also known as West Malaysia or the Malaysian Peninsula, ...
,
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
,
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
, Cambodia,
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
,
Hainan Hainan (, ; ) is the smallest and southernmost province of the People's Republic of China (PRC), consisting of various islands in the South China Sea. , the largest and most populous island in China,The island of Taiwan, which is slightly l ...
,
South-Central China South Central China, South-Central China or Central-South China ( zh, c = 中南, p = Zhōngnán, l = Central-South), is a region of the People's Republic of China defined by State Council that includes the provinces of Guangdong, Hainan, He ...
including
Yunnan Yunnan , () is a landlocked Provinces of China, province in Southwest China, the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is ...
,
Andaman and Nicobar Islands The Andaman and Nicobar Islands is a union territory of India consisting of 572 islands, of which 37 are inhabited, at the junction of the Bay of Bengal and the Andaman Sea. The territory is about north of Aceh in Indonesia and separated f ...
,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
,
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...
,
Assam Assam (; ) is a state in northeastern India, south of the eastern Himalayas along the Brahmaputra and Barak River valleys. Assam covers an area of . The state is bordered by Bhutan and Arunachal Pradesh to the north; Nagaland and Manipur ...
,
East Himalaya ] The Eastern Himalayas extend from eastern Nepal across Northeast India, Bhutan, the Tibet Autonomous Region to Yunnan in China and northern Myanmar. The climate of this region is influenced by the monsoon of South Asia from June to September. It ...
,
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainous ...
,
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is mai ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
(from
Punjab Punjab (; Punjabi: پنجاب ; ਪੰਜਾਬ ; ; also romanised as ''Panjāb'' or ''Panj-Āb'') is a geopolitical, cultural, and historical region in South Asia, specifically in the northern part of the Indian subcontinent, comprising ...
and
Gujarat Gujarat (, ) is a state along the western coast of India. Its coastline of about is the longest in the country, most of which lies on the Kathiawar peninsula. Gujarat is the fifth-largest Indian state by area, covering some ; and the ninth ...
to
Assam Assam (; ) is a state in northeastern India, south of the eastern Himalayas along the Brahmaputra and Barak River valleys. Assam covers an area of . The state is bordered by Bhutan and Arunachal Pradesh to the north; Nagaland and Manipur ...
,
Mizoram Mizoram () is a state in Northeast India, with Aizawl as its seat of government and capital city. The name of the state is derived from "Mizo people, Mizo", the endonym, self-described name of the native inhabitants, and "Ram", which in the Mizo ...
,
Maharashtra Maharashtra (; , abbr. MH or Maha) is a states and union territories of India, state in the western India, western peninsular region of India occupying a substantial portion of the Deccan Plateau. Maharashtra is the List of states and union te ...
,
West Bengal West Bengal (, Bengali: ''Poshchim Bongo'', , abbr. WB) is a state in the eastern portion of India. It is situated along the Bay of Bengal, along with a population of over 91 million inhabitants within an area of . West Bengal is the fourt ...
and South India), and
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
.


Habitat

''Dillenia pentagyna'' is found in rainforests, thickets and on hills below 400m. In Cambodia it grows in groups in open/clear forests. In the ''
Pinus merkusii ''Pinus merkusii'', the Merkus pine or Sumatran pine, is a pine native to the Malesia region of southeast Asia, and the only one that occurs naturally south of the equator. Description ''Pinus merkusii'' is a medium-sized to large tree, reachin ...
''-dominated forests of
Kirirom National Park The Kirirom National Park ( km, ឧទ្យានជាតិគិរីរម្យ), officially the Preah Suramarit-Kossamak Kirirom National Park ( km, ឧទ្យានជាតិព្រះសុរាម្រិត-កុសមៈ គ ...
, southwestern Cambodia (which occur from 400-1000m elevation), the tree occurs in the 2-8m high understorey under the 20m high pine canopy. ''D. pentagyna'' grows on islands in the Mekong in Steung Treng and Kratie provinces, northeastern Cambodia. It is found in the ''Deciduous, dipterocarp, seasonal, hardwood forest'', dominated by
Dipterocarpaceae Dipterocarpaceae is a family of 16 genera and about 695 known species of mainly tropical lowland rainforest trees. The family name, from the type genus ''Dipterocarpus'', is derived from Greek (''di'' = two, ''pteron'' = wing and ''karpos'' = fru ...
species, occurring above metamorphic sandstone bedrock, at 25-30m elevation. In
Phnom Kulen National Park The Phnom Kulen National Park ( km, ឧទ្យានជាតិភ្នំគូលែន) is a national park in Cambodia, located in the Phnom Kulen mountain massif in Siem Reap Province. It was established in 1993 and covers . Its official ...
, Siem Reap Province, northwestern Cambodia, the tree is a common canopy species in the Deciduous Dipterocarp Forest.


Uses

The tree is used as a source of fruit, traditional medicine, timber and charcoal. Amongst the
Tetun , nativename=Tetun , states= Indonesia East Timor , speakers=, mostly in Indonesia , date=2010–2011 , ref=e18 , speakers2=50,000 L2-speakers in Indonesia and East Timor , familycolor=Austronesian , fam2=Malayo-Polynesian , fam3= Central–East ...
speaking practitioners of the ''ai tahan'' traditional medicine, in
Belu Regency Belu Regency is a regency in East Nusa Tenggara province of Indonesia. Established on 20 December 1958, Belu Regency has its seat (capital) in the large town of Atambua. In December 2012 a separate Regency - Malaka Regency - was created from th ...
, Timor Barat, Indonesia, the bark is used to treat headaches and migraines. In Cambodia, the fruit is eaten, but is not generally appreciated, it also provides a component for a remedy against cough. The timber is used to make humidity resistant boards and beams. Small knick-knacks are made from the wood, which makes excellent charcoal as well. The Bunong people of Mondulkiri Province, northeastern Cambodia, drink a decoction of the bark and wood of both this tree and ''
Oroxylum indicum ''Oroxylum indicum'' is a species of flowering plant belonging to the monotypic genus ''Oroxylum'' and the family Bignoniaceae, and is commonly called Indian trumpet tree, oroxylum, Indian trumpet flower, broken bones, Indian caper, scythe tree o ...
'' to treat cold and fever. Amongst
Kuy Kuy, also known as Kui, Suay or Kuay ( th, ภาษากูย; km, ភាសាកួយ), is a Katuic languages, Katuic language, part of the larger Austroasiatic languages, Austroasiatic family spoken by the Kuy people of Southeast Asia. ...
- and Khmer-speaking people living in the same villages in Stung Treng and Preah Vihear provinces of north-central Cambodia, the tree is used as source of medicine, fuel and food. In various areas of India, parts of the plant are used in ethnomedicine. The Koch-Rajbanshi people of western Assam use the seed and bark against cancer. In the
Konkan The Konkan ( kok, कोंकण) or Kokan () is a stretch of land by the western coast of India, running from Damaon in the north to Karwar in the south; with the Arabian Sea to the west and the Deccan plateau in the east. The hinterland ...
region of Maharashtra, village people use a paste of water and bark to treat wounds. In Deogarh district, Tribal people use a decoction of the fruit and '' Zingiber montanum'' for "blood dysentery". To treat diabetes, ripe fruit is taken regularly. The unripe fruit is consumed as a vegetable.


References

{{Taxonbar, from=Q10960107 pentagyna Flora of South-Central China Flora of Hainan Flora of the Indian subcontinent Flora of Indo-China Flora of Java Flora of the Lesser Sunda Islands Flora of Malaya Flora of Sulawesi Plants described in 1795