''Dennstaedtia punctilobula'', the eastern hayscented fern or hay-scented fern, is a species of

fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...

native to eastern

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

, from

Newfoundland Newfoundland and Labrador (; french: Terre-Neuve-et-Labrador; frequently abbreviated as NL) is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region ...

west to

Wisconsin Wisconsin () is a state in the upper Midwestern United States. Wisconsin is the 25th-largest state by total area and the 20th-most populous. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake M ...

and

Arkansas Arkansas ( ) is a landlocked state in the South Central United States. It is bordered by Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, and Texas and Oklahoma to the west. Its name is from the Osage ...

, and south in the

Appalachian Mountains The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. They ...

to northern

Alabama (We dare defend our rights) , anthem = "Alabama (state song), Alabama" , image_map = Alabama in United States.svg , seat = Montgomery, Alabama, Montgomery , LargestCity = Huntsville, Alabama, Huntsville , LargestCounty = Baldwin County, Al ...

; it is most abundant in the east of its range, with only scattered populations in the west. It is a

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...

fern with

frond A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the lar ...

s growing to 40–100 cm (rarely 130 cm) tall and 10–30 cm broad; the fronds are bipinnate, with pinnatifid pinnules about three times as long as broad. It occurs in damp or dry acidic soils in woods or open woods, from sea level up to 1,200 m altitude. ''Dennstaedtia punctilobula'' can exhibit varying degrees of

phototropism Phototropism is the growth of an organism in response to a light stimulus. Phototropism is most often observed in plants, but can also occur in other organisms such as fungi. The cells on the plant that are farthest from the light contain a hor ...

. The

common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...

"Hay-scented Fern" comes from the fact that crushing it produces an aroma of fresh

hay Hay is grass, legumes, or other herbaceous plants that have been cut and dried to be stored for use as animal fodder, either for large grazing animals raised as livestock, such as cattle, horses, goats, and sheep, or for smaller domesticat ...

. The presence of ''Dennstaedtia punctilobula'' influences the dynamics of the

understory In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abov ...

vegetation of many forests in the eastern United States. An abundance of ''

Rubus allegheniensis ''Rubus allegheniensis'' is a North American species of glandular highbush blackberry in section Alleghenienses of the genus ''Rubus'', a member of the rose family. It is the most common and widespread highbush blackberry in eastern and centra ...

'' in open areas encourages new tree seedlings. Where the effects of herbivorous animals (such as

deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...

) reduce the abundance of ''Rubus allegheniensis'', ''Dennstaedtia punctilobula'', which is not browsed by deer, takes over. Where ''Dennstaedtia punctilobula'' becomes common, the growth of tree seedlings is restricted.

References

{{Taxonbar, from=Q2863582 punctilobula Ferns of the Americas Ferns of Canada Ferns of the United States Flora of Eastern Canada Flora of the Northeastern United States Flora of the Southeastern United States Flora of the Appalachian Mountains Flora of the Great Lakes region (North America) Plants described in 1803 Least concern flora of the United States Least concern biota of North America Garden plants of North America