Dendrochirus Biocellatus
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''Dendrochirus biocellatus'', the twospot turkeyfish, twinspot lionfish, twoeyed lionfish or ocellated lionfish, is a species of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...
belonging to the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Scorpaenidae The Scorpaenidae (also known as scorpionfish) are a family of mostly marine fish that includes many of the world's most venomous species. As their name suggests, scorpionfish have a type of "sting" in the form of sharp spines coated with venom ...
, the scorpionfishes and lionfishes. This species is widespread throughout the
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
waters of the
Indo-West Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
region, In the wild, the species eats small fish as well as shrimp.


Taxonomy

''Dendrochirus biocellatus'' Was first formally described as ''Nemapterois biocellatus'' in 1938 by the American
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...
Henry Weed Fowler Henry Weed Fowler (March 23, 1878 – June 21, 1965) was an American zoologist born in Holmesburg, Pennsylvania. He studied at Stanford University under David Starr Jordan. He joined the Academy of Natural Sciences in Philadelphia and worked as ...
with the type locality given as off Jolo Light on
Jolo Island Jolo ( tsg, Sūg) is a volcanic island in the southwest Philippines and the primary island of the province of Sulu, on which the capital of the same name is situated. It is located in the Sulu Archipelago, between Borneo and Mindanao, and has ...
in the
Sulu Sea The Sulu Sea ( fil, Dagat Sulu; Tausug: ''Dagat sin Sūg''; Chavacano: ''Mar de Sulu''; Cebuano: ''Dagat sa Sulu''; Hiligaynon: ''Dagat sang Sulu''; Karay-a: ''Dagat kang Sulu''; Cuyonon: ''Dagat i'ang Sulu''; ms, Laut Sulu) is a body o ...
in the Philippines. This is the only species in the
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
''Nemapterois'', this name being a compound of ''nema'', meaning “thread” and ''
Pterois ''Pterois'' is a genus of venomous marine fish, commonly known as lionfish, native to the Indo-Pacific. Also called firefish, turkeyfish, tastyfish, or butterfly-cod, it is characterized by conspicuous warning coloration with red, white, cre ...
'', the genus of “typical” lionfishes. The specific name biocellatus means two spots, alluding to the two eye like spots, called ocelli, on the soft rayed part of the
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
.


Description

''Dendrochirus biocellatus'' has 13 spines and 9 soft rays in its dorsal fin with 3 spines and 5 soft rays in the
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
, the middle spines in the dorsal fin are shorter than the depth of the body, with deep incisions in the fin membranes of that fin too. The large, wing-like
pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
s have 20 or 21 fin rays with the upper rays fully connected by membranes, the lower rays being simple and their tips extend beyond the membrane. The tentacles on the lachrimal bone are long, their length being more than two times the diameter of the eye. The overall colour of this lionfish is reddish-brown. there are 2, sometimes 3, large black eye like spots, or ocelli, on the soft-rayed part of the dorsal fin, there are 3 pink to yellowish bars on the flanks. The pectoral fins are banded with pale and brown. This species attains a maximum
total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...
of .


Distribution and habitat

''Dendrochirus biocellatus'' has a wide
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
distribution which extends from the western
Mascarene Islands The Mascarene Islands (, ) or Mascarenes or Mascarenhas Archipelago is a group of islands in the Indian Ocean east of Madagascar consisting of the islands belonging to the Republic of Mauritius as well as the French department of Réunion. Thei ...
, The Maldives and Sri Lanka eastwards to the
Mariana Mariana may refer to: Literature * ''Mariana'' (Dickens novel), a 1940 novel by Monica Dickens * ''Mariana'' (poem), a poem by Alfred Tennyson, 1st Baron Tennyson * ''Mariana'' (Vaz novel), a 1997 novel by Katherine Vaz Music *"Mariana", a so ...
and
Tuamotu Islands The Tuamotu Archipelago or the Tuamotu Islands (french: Îles Tuamotu, officially ) are a French Polynesian chain of just under 80 islands and atolls in the southern Pacific Ocean. They constitute the largest chain of atolls in the world, extendin ...
, north as far as southern Japan, south to
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
and Tonga. In Australian waters it is found at Scott Reef off
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
, at the
Ashmore Ashmore is a village and civil parish in the North Dorset district of Dorset, England, southwest of Salisbury. The village is centred on a circular pond and has a church and several stone cottages and farms, many with thatched roofs. It is ...
and Hibernia reefs in the
Timor Sea The Timor Sea ( id, Laut Timor, pt, Mar de Timor, tet, Tasi Mane or ) is a relatively shallow sea bounded to the north by the island of Timor, to the east by the Arafura Sea, and to the south by Australia. The sea contains a number of reefs, ...
and at
Christmas Island Christmas Island, officially the Territory of Christmas Island, is an Australian external territory comprising the island of the same name. It is located in the Indian Ocean, around south of Java and Sumatra and around north-west of the ...
. This species is found at depths of in area where there is rich coral growth on reefs in clear water.


Biology

''Dendrochirus biocellatus'' shelters in caves and on ledges during the day, emerging at night to feed on small fishes and small crustaceans. The fin spines are
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
ous. This species has been observed to have unusual feeding behaviour, as it approaches prey it snaps its dorsal fin spines and shakes its head from side-to-side. This be designed to distract, or possibly even attract, the prey to the fish. It has also been suggested that the fleshy tentacles on the upper jaw, which give it the name of "Fu-Manchu lionfish" in the
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
trade, may be used to attract prey to within striking distance. These fish stalk prey by moving stealthily along the substrate or around reef structures, moving forward, either by “hopping” on the
pelvic fins Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two en ...
or by sinuously waving its
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
until it is around half its body length from its quarry, when it makes a rapid lunge and ingests the prey.


Utilisation

''Dendrochirus biocellatus'' appears occasionally in the aquarium trade. They are considered to be moderately difficult to keep in aquaria and not for beginners.


References

* *


External links

* {{Taxonbar, from=Q2371681 biocellatus Fish described in 1938 Taxa named by Henry Weed Fowler