Dendrobium Tetragonum
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''Dendrobium tetragonum'', commonly known as the tree spider orchid, is a variable species of
epiphytic An epiphyte is an organism that grows on the surface of a plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphytes grow are called phoroph ...
or
lithophytic Lithophytes are plants that grow in or on rocks. They can be classified as either epilithic (or epipetric) or endolithic; epilithic lithophytes grow on the surfaces of rocks, while endolithic lithophytes grow in the crevices of rocks (and are als ...
orchid Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Along with the Asteraceae, they are one of the two largest families of flowering ...
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to eastern
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
. Tree spider orchids are unusual in having pendulous
pseudobulb The pseudobulb is a storage organ found in many epiphytic and terrestrial sympodial orchids. It is derived from a thickening of the part of a stem between leaf nodes and may be composed of just one internode or several, termed heteroblastic and ...
s that are thin and wiry near the base then expand into a fleshy, four-sided upper section before tapering at the tip. There are only a few thin but leathery leaves at the end of the pseudobulbs and up to five flowers on relatively short flowering stems. To allow for the variations in the species there are five subspecies and a variety, some with a unique common name.


Description

''Dendrobium tetragonium'' is an epiphytic or lithophytic
herb In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables and other plants consumed for macronutrients, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal ...
with pendulous pseudobulbs long and wide. The pseudobulbs have a thin, wiry base but expand to a fleshy, four-sided upper section and a tapering tip with between two and five thin but leathery leaves. The leaves are long and wide. The flowering stems are long and bear up to eight flowers. The number, size and colour of the flowers depend on subspecies. Flower size ranges from long and wide. The flowers range in colour from green to pale yellow, sometimes with red blotches and a labellum that ranges in colour from white, to yellowish with red, brown or purplish markings. The
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
s range in size from long and wide and the
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s from long and wide. The labellum ranges from long and from wide. Flowering occurs between April and November.


Taxonomy and naming

''Dendrobium tetragonum'' was first formally described in 1839 by Allan Cunningham from a specimen "hanging loosely from the stems of small trees in ''dry'' shaded woods,
Moreton Bay Moreton Bay is a bay located on the eastern coast of Australia from central Brisbane, Queensland. It is one of Queensland's most important coastal resources. The waters of Moreton Bay are a popular destination for recreational anglers and are ...
". The description was published in '' Edwards's Botanical Register'' edited by
John Lindley John Lindley FRS (5 February 1799 – 1 November 1865) was an English botanist, gardener and orchidologist. Early years Born in Catton, near Norwich, England, John Lindley was one of four children of George and Mary Lindley. George Lindley w ...
.


Subdivisions

* ''Dendrobium tetragonum'' var. ''cacatua'' (
D.L.Jones David Lloyd Jones (born 1944) is an Australians, Australian horticultural botanist and the author of many books and papers, especially on Australian orchids. Jones was born in Victoria (Australia), Victoria and in his youth was a student at Burnl ...
& M.A.Clem.) H.Mohr
– the yellow tree spider orchid has greenish to pale yellow flowers with a few fine reddish purple markings and a white labellum, usually with no markings, and flowers from July to September; * ''Dendrobium tetragonum'' subsp. ''cataractarum'' Peter B.Adams, S.D.Lawson & G.A.Peterson has yellowish green flowers with red, purple or brown marks on the edge of the sepals and a pale, cream-coloured labellum with reddish purple markings, and flowers from August to September; * ''Dendrobium tetragonum'' subsp. ''giganteum'' ( Leaney) Peter B.Adams – the blotched tree spider orchid has greenish yellow flowers with prominent red blotches and a white labellum with red and purplish lines, and flowers from April to November; * ''Dendrobium tetragonum'' subsp. ''melaleucaphilum'' (
D.L.Jones David Lloyd Jones (born 1944) is an Australians, Australian horticultural botanist and the author of many books and papers, especially on Australian orchids. Jones was born in Victoria (Australia), Victoria and in his youth was a student at Burnl ...
& M.A.Clem.) Dockrill
– the flared tree spider orchid has the largest flowers in the species, the flowers green to greenish yellow with reddish blotches and a white or pale yellow labellum with red or purplish marks, and flowers from May to October; * ''Dendrobium tetragonum'' subsp. ''serpentis'' Peter B.Adams has flowers that are green at first, then turn yellow with few or no markings apart from on the labellum which is white to pale cream-coloured with reddish purple markings, and flowers from August to September; * ''Dendrobium tetragonum'' subsp. ''tetragonum'' – the banded tree spider orchid has the smallest flowers of the species, the flowers green to greenish yellow with dark reddish brown bands on the edges and a cream-coloured or yellow labellum with brown, red or purplish marks, and flowers from May to October.


Distribution and habitat

Tree spider orchids are usually epiphytes which grow on trees in rainforest or shady places beside streams, but also sometimes on
paperbark ''Melaleuca'' () is a genus of nearly 300 species of plants in the myrtle family, Myrtaceae, commonly known as paperbarks, honey-myrtles or tea-trees (although the last name is also applied to species of ''Leptospermum''). They range in size f ...
trees, especially ''
Melaleuca styphelioides ''Melaleuca styphelioides'', known as the prickly-leaved paperbark or prickly paperbark, is a plant native to eastern Australia. It is a tree with spongy bark, prickly leaves and spikes of creamy-white flowers. Description It is a small to medi ...
'' and occasionally on rocks. Different forms of the species occur in different areas of coastal or near-coastal
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
and
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
. * ''Dendrobium tetragonum'' var. ''cacatua'' is found from the Mount Windsor Tableland near
Cape Tribulation Cape Tribulation is a headland and coastal locality in the Shire of Douglas in northern Queensland, Australia. In the , Cape Tribulation had a population of 118 people. Geography The locality is north of Cairns. It is within the Daintree Na ...
to the
Eungella National Park Eungella National Park ( ; meaning "Land of the clouds") is a protected area in Queensland, Australia. It is on the Clarke Range at the end of the Pioneer Valley 80 km west of Mackay, and 858 km northwest of Brisbane. Eungel ...
in Queensland; * Subspecies ''cataractarum'' is restricted to Connors Range west of the Cape Palmerston National Park in central eastern Queensland; * ''Dendrobium tetragonum'' subsp. ''giganteum'' occurs between the
Iron Range The term Iron Range refers collectively or individually to a number of elongated iron-ore mining districts around Lake Superior in the United States and Canada. Much of the ore-bearing region lies alongside the range of granite hills formed by ...
and
Mackay Mackay may refer to: *Clan Mackay, the Scottish clan from which the surname "MacKay" derives Mackay may also refer to: Places Australia * Mackay Region, a local government area ** Mackay, Queensland, a city in the above region *** Mackay Airpor ...
in Queensland; * ''Dendrobium tetragonum'' subsp. ''melaleucaphilum'' is found from the
Blackdown Tableland National Park Blackdown Tableland is a national park in the Central Highlands Region, Queensland, Australia. Geography The park is in Central Queensland, northwest of Brisbane. The mountainous terrain of the tablelands provides a unique landscape featurin ...
in Queensland to the Blue Mountains in New South Wales; * ''Dendrobium tetragonum'' subsp. ''serpentis'' is restricted to the northern parts of the Blackdown Tableland; * ''Dendrobium tetragonum'' subsp. ''tetragonum'' ranges from
Fraser Island Fraser Island (Butchulla: ) is a World Heritage-listed island along the south-eastern coast in the Wide Bay–Burnett region, Queensland, Australia. The island is approximately north of the state capital, Brisbane, and is within the Fraser ...
in Queensland to
Tomerong Tomerong is a village in the South Coast region of New South Wales, Australia. It is located approximately 180 kilometres by road south from the state capital Sydney and 20 kilometres south of the commercial centre of the City of Shoalhaven, Nowr ...
in New South Wales.


References


External links


Atlas of Living Australia, ''Dendrobium tetragonum'' A.Cunn. Tree Spider OrchidAndy's Orchids (Encinitas California): ''Dendrobium tetragonum'' subsp. ''giganteum''
{{DEFAULTSORT:Dendrobium Tetragonum tetragonum Endemic orchids of Australia Orchids of New South Wales Orchids of Queensland Plants described in 1839