''Demersatheca'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of extinct

vascular plant Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They al ...

s of the

Early Devonian The Early Devonian is the first of three Epoch (geology), epochs comprising the Devonian period, corresponding to the Lower Devonian Series (stratigraphy), series. It lasted from and began with the Lochkovian Stage , which was followed by the P ...

(Pragian, around ). Fossils were first found in the Posongchong Formation of eastern Yunnan, China. The plant had smooth leafless stems at least 1 mm in diameter, but only regions which bore

spore In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, f ...

-forming organs or

sporangia A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point in their life cy ...

are well-known. Sporangia were borne in 'spikes' or strobili, at least 40 mm long; one had 32 sporangia. Sporangia were arranged in four rows, two sporangia being opposite to one another on the stem with the next two being at right angles. Each sporangium consisted of two 'valves' which opened at the top to release their spores. A particular feature of ''Demersatheca'' which distinguishes it from other

zosterophyll The zosterophylls are a group of extinct land plants that first appeared in the Silurian period. The taxon was first established by Banks in 1968 as the subdivision Zosterophyllophytina; they have since also been treated as the division Zosteroph ...

s is that the stalk-less sporangia were sunken into the stem of the spike, so that the outer valve was flush with the surface.

Taxonomy

''D. contigua'' was initially called ''

Zosterophyllum ''Zosterophyllum'' was a genus of Silurian-Devonian vascular land plants with naked branching axes on which usually kidney-shaped sporangia were arranged in lateral positions. It is the type genus for the group known as zosterophylls, thought to ...

contiguum'' by Li and Cai, based on parts of two spikes (strobili) of sporangia. When further specimens were found, a review by Li and Edwards agreed that the plant was a zosterophyll, but decided that the sunken sporangia and the absence of sporangial stalks were sufficient to require a new genus, ''Demersatheca''. The name is derived from ''demersus'', Latin for sunken, and ''theca'', Latin for container, referring to the organization of the sporangia. Hao and Xue in 2013 considered the genus as ''

incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...

'' within the tracheophytes (vascular plants).

References

{{Taxonbar, from=Q5255175 Early Devonian plants Prehistoric plant genera