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The excitable delma (''Delma tincta'') is a species of lizard in the
Pygopodidae Pygopodidae, commonly known as legless lizards, snake-lizards, or flap-footed lizards, is a family of squamates with reduced or absent limbs, and are a type of gecko. At least 35 species are placed in two subfamilies and eight genera. They hav ...
family
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to Australia. The lizard gets its name from its active and jumpy defense mechanism. It will erratically jump multiple times in succession, each jump pivoting its body in a different direction. ''D. tincta'' is a slender, long legless lizard that through evolution lost its limbs. It is found throughout Australia in a variety of habitats, and spends most of its time hiding. Due to its nocturnal nature, it is rare to spot in the wild. This legless lizard is small to moderate in size, with a tail that is three to four times its body length. The typical size of the excitable delma is 250 – 300 mm. This lizard is an insectivore and feeds on insects it finds when travelling through grass, logs, surface soil, and loose rocks. Like all pygopodids, the excitable delma is oviparous and only lays two eggs per clutch.


Taxonomy

The excitable delma is officially classified as ''Delma tincta'' from 1888 by De Vis, but in 1901, Garman referred to it as ''Delma reticulata''. The twelve subgenera are: * Marble-faced delma ''Delma australis'' * Rusty-topped delma ''Delma borea'' * Pilbara delma ''Delma elegans'' * Fraser's delma ''Delma fraseri'' * ''
Delma grayii ''Delma grayii'', also known as side-barred delma or Gray's legless lizard, is a species of lizard in the Pygopodidae family endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, ...
'' * Striped legless lizard ''Delma impar'' * Patternless delma ''Delma inornata'' * ''
Delma molleri ''Delma molleri'' (commonly known as Gulfs delma and Adelaide delma) is a small he snout-vent length of the largest specimen measured limbless lizard found in southern South Australia around the Adelaide Hills, and pretty common in Adelaide s ...
'' * ''
Delma nasuta ''Delma nasuta'', also known as sharp-snouted delma or sharp-snouted legless lizard, is a species of lizard in the Pygopodidae family endemic Endemism is the state of a species being found in a single defined geographic location, such as a ...
'' * Peace delma ''Delma pax'' * Leaden delma ''Delma plebeia'' * Collared delma ''Delma torquata''


Description

The excitable delma looks remarkably like a
snake Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more j ...
, because through
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
, the species has completely lost its limbs. It has an extremely long and slender body with a tail that is three to four times its body length. The excitable delma's snout is also rounded in shape. The body size from snout to vent in an adult excitable delma is about 80 to 90 mm, with a tail that extends 240 mm. Juveniles' measurements are typically 40 mm for the body and 136 mm for the tail. They are small to moderate in size compared to the rest of the ''Delma'' genus. The body of the excitable delma is an olive green to grey-brown colour with a white cream underbody, and consists of 14 rows of midbody scales, three subocular supralabial scales, and one pair of supranasal scales. It has three distinct black markings across the upper section of the head and neck. The black markings are intersected by four yellow bands, the first band on the muzzle, the second before the eye, the third behind the eye, and the final across the back part of the head. The markings on the excitable delma's head and neck are more dominant on the juveniles.


Location

Originally, the excitable delma was believed to only be located in
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , establishe ...
and north-eastern
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
. Studies have now found this statement to be incorrect, and this legless lizard has been observed in five of the eight states and territories of Australia. The distribution of the lizard is: * Queensland - widespread throughout * New South Wales - northern sections * Northern Territory - southern two thirds * South Australia - north-eastern corner extending across to the south-west * Western Australia - central west to the north An
observational study In fields such as epidemiology, social sciences, psychology and statistics, an observational study draws inferences from a sample to a population where the independent variable is not under the control of the researcher because of ethical concer ...
by Glenn M. Shea in 1991 resulted in 10 specimens of excitable delma being reported around Australia. Seven specimens were examined in Queensland, one in New South Wales, one in a museum in
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
, and the last observed from Mt. Barker, which could be the one close to either Adelaide,
South Australia South Australia (commonly abbreviated as SA) is a state in the southern central part of Australia. It covers some of the most arid parts of the country. With a total land area of , it is the fourth-largest of Australia's states and territories ...
, or Perth, Western Australia. The excitable delma has a wide variety of habitats from
dry forests The tropical and subtropical dry broadleaf forest is a habitat type defined by the World Wide Fund for Nature and is located at tropical and subtropical latitudes. Though these forests occur in climates that are warm year-round, and may receive ...
and spinifex deserts to rocky outcrops. They are usually found under loose rocks, logs or similar debris, and sometimes surface layers of
soil Soil, also commonly referred to as earth or dirt Dirt is an unclean matter, especially when in contact with a person's clothes, skin, or possessions. In such cases, they are said to become dirty. Common types of dirt include: * Debri ...
.


Ecology

The excitable delma gets its name because it is the most active species in the pygopodids. It has a unique jumping
defense mechanism In psychoanalytic theory, a defence mechanism (American English: defense mechanism), is an unconscious psychological operation that functions to protect a person from anxiety-producing thoughts and feelings related to internal conflicts and o ...
in which its body twists in a display of acrobatics to scare or confuse predators. When the excitable delma is agitated, it uses its long tail to generate an upward thrust, which causes its entire body to lift from the ground. When the delma leaps it erratically changes its direction with each jump. The
University of California, Berkeley The University of California, Berkeley (UC Berkeley, Berkeley, Cal, or California) is a public land-grant research university in Berkeley, California. Established in 1868 as the University of California, it is the state's first land-grant u ...
observed an adult excitable delma in captivity; without any stimulus, the delma was filmed jumping more than 30 times. The jumps reached heights of 11 to 15 cm and lasted 0.41 to 0.45 seconds from initial lift to impact. The excitable delma moves through the different ecosystems it inhabits in a locomotor motion known as saltation. Saltation is common in limbless vertebrates, and in the pygopodids family is a behavioral response to
predation Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
. It can be sometimes be confused as an
escape response Escape response, escape reaction, or escape behavior is a mechanism by which animals avoid potential predation. It consists of a rapid sequence of movements, or lack of movement, that position the animal in such a way that allows it to hide, freeze ...
, but this is not the function of this behaviour. Saltation is the locomotion motion of side-pushing, and the jumping in delmas allows them to move their bodies at a greater wave-speed. The locomotion movement begins in the tail, and the lizard has evolved to propel itself forward at a 45° angle. Another defense mechanism of the species is the capability to vocalize sounds like other lizards in the pygopodids.


References

{{Taxonbar, from=Q5262749 Pygopodids of Australia Delma Reptiles described in 1888 Endemic fauna of Australia