Deer Brush
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''Ceanothus integerrimus'', known by the common name deer brush, is a species of woody
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
in the family
Rhamnaceae The Rhamnaceae are a large family of flowering plants, mostly trees, shrubs, and some vines, commonly called the buckthorn family. Rhamnaceae is included in the order Rosales. The family contains about 55 genera and 950 species. The Rhamnaceae h ...
, native to the western United States in Arizona, New Mexico, California, Oregon, and Washington. It grows in montane chaparral and woodlands regions, in hardwood forests, and in fir, spruce, and Ponderosa pine plant communities, being most abundant in the
California chaparral and woodlands The California chaparral and woodlands is a terrestrial ecoregion of southwestern Oregon, northern, central, and southern California (United States) and northwestern Baja California (Mexico), located on the west coast of North America. It is a ...
and
Sierra Nevada The Sierra Nevada () is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primarily ...
. akley/ref> epson/ref>


Description

''Ceanothus integerrimus'' is a deciduous shrub from tall with an open ascending to erect branch habit. epson/ref> It is a drought-tolerant phanerophyte. Nitrogen-fixing actinomycete bacteria form root nodules on ''Ceanothus'' roots. oward/ref> Its stems are round yellow to pale green in color with either small soft to straight stiff sharp hairs parallel to or in contact with the surface of the stem,. epson/ref> The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are glossy, deciduous and 2.5–8 cm long. Leaves grow alternately on stems. The leaf petioles are less than 15 mm in length and the stipules are also deciduous. The leaf blade is lanceolate, elliptical or oblong to widely ovate in shape. Leaves can have one to two ribs from the base; they are also generally thin and have an acute to obtuse tip. Leaf margins are either entire or slightly dentate, more so towards the leaf tip. Leaf surfaces are light green and are ciliate or contain hairs visible only by magnification. The lower leaves are also hairy and lighter in color. epson/ref> The flowers are white or blue and rarely pink in color. They are produced in raceme clusters of 15 centimeters or less and contain both male and female organs. The fruit is a sticky valved capsule about 4–5 mm in diameter with a slight crest; the seed is ejected from the capsule after splitting. epson/ref> It regenerates by seed, shoot formation from the crown and stem, and also by layering when branches come in contact with soil. oward/ref> riffin/ref> It has been suggested that some ''Ceanothus'' species do not resprout from the root after the crown has burned as a result of fire where most other species are able to regenerate. Pollination of flowers is primarily by
bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyly, monophyletic lineage within the ...
s. Seed production occurs after about four years of age. High densities of seeds occur in the upper soil of ''Ceanothus'' communities. Seeds remain viable up to 24 years or more. Seed dormancy is broken by the removal of the seed coat by fire scarification or physical disturbance. oward/ref> riffin/ref> Seeds germinate best at about 1 inch soil depth in shady areas in the spring following fire scarification. oward/ref> riffin/ref>


Varieties

There are four weakly defined varieties of ''Ceanothus integerrimus''. Identification is primarily by leaf morphology and flower color: epson/ref> oward/ref> riffin/ref> *''Ceanothus integerrimus'' var. ''californicus''. Leaves elliptic, lanceolate or oblong to ovate in shape and are three ribbed, from the leaf base. Leaf surfaces have small hairs and the undersides are less hairy than the surface. Flowers generally white or blue. *''Ceanothus integerrimus'' var. ''integerrimus''. *''Ceanothus integerrimus'' var. ''macrothyrsus''. Leaf blades oblong or ovate. Leaf bases are three ribbed at the base. Leaf surfaces are pubescent on both the surface and undersides. Flowers are white. *''Ceanothus integerrimus'' var. ''puberulus''. Leaf blades elliptical or lanceolate and oblong to obovate in shape. Leaf base is three ribbed from the leaf base. Leaves are also pubescent on both sides. Flowers white. ''Ceanothus integerrimus'' hybridizes with ''Ceanothus tomentosus'' (Lemmon's ceanothus) and ''Ceanothus cordulatus'' (mountain whitethorn).


Ecology

''C. integerrimus'' is an important part of forest regeneration after wildfires by providing nitrogen. It does this by creating nitrogen rich patches in the soil. The nitrogen source is created by its root association with nitrogen fixing bacteria. Deer and specifically mule deer feed on ''C. integerrimus''.
Porcupine Porcupines are large rodents with coats of sharp spines, or quills, that protect them against predation. The term covers two families of animals: the Old World porcupines of family Hystricidae, and the New World porcupines of family, Erethizont ...
s and quail have also been observed eating the stems and seeds. Nutritionally leaves are a good source of protein and stems and leaves also contain high levels of calcium. However, nutritional quality of leaves is seasonal and appears to be best from fall to early spring. oward/ref>


Uses

Indigenous peoples of California use the branches to treat women after childbirth. The Miwok Indians of
Northern California Northern California (colloquially known as NorCal) is a geographic and cultural region that generally comprises the northern portion of the U.S. state of California. Spanning the state's northernmost 48 counties, its main population centers incl ...
also use the branches of this plant in weaving complex baskets. oward/ref> The Concow tribe call the tree hē′-bē ( Konkow language). Cattle ranchers will claim that cattle do immensely well on this plant during their seasonal grazing.


References


Bibliography

# Debano, L. F. & Conrad, C. E. (1978). The Effect of Fire on Nutrients in the Chaparral Ecosystem. ''Ecology'' 59 (3): 489–497. # Gibbens, R. R. P. & Schultz, A. M. (1963). Brush manipulation on a deer winter range. ''California Fish and Game'' 49 (2): 95–118. 976 # Griffin, James R. (1982). Pine seedlings, native ground cover, and Lolium multiflorum on Marble-Cone burn, Santa Lucia Range, CA. ''Madrono'' 29 (3): 177–188.
Jepson Flora Treatment: ''Ceanothus integerrimus''
# Howard, Janet L. (1997). System ''Ceanothus integerrimus'' In: Fire Effects Information nline U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory

# Munz, P. A. & Keck, D. D. (1959). ''A California Flora with Supplement'' p. 973. University of California Press, Berkeley, Los Angeles. # Moerman, D. ''Native American Ethnobotany''. Timber Press, Oregon: 1988. # Oakley B. B., North, M. P., & Franklin, J. F. (2003). The effects of fire on soil nitrogen associated with patches of the actinorhizal shrub ''Ceanothus cordulatus''. ''Plant and Soil'' 254: 35–46. # Oswald, V. H. & Ahart, L. (1994). ''Manual of the Vascular Plants of Butte County, California'' p. 192. Native Plant Society, Sacramento. # Raven, P. H. & Axelrod, D. I. (1977). ''Origin and relationships of the California Flora''. University of California Publications in Botany 72. Sacramento: University of California Press. # Russell, C. P. (1932). Seasonal Migration of Mule Deer. ''Ecological Monographs'' 2:1 pp. 1–46.


External links


USDA Plant Profiles: ''Ceanothus integerrimus''Jepson Interchange – ''Ceanothus integerrimus'''Ceanothus integerrimus' – Photo Gallery
{{Taxonbar, from=Q5055724 Ceanothus, integerrimus Flora of Washington (state) Flora of Oregon Flora of Arizona Flora of New Mexico Flora of the Sierra Nevada (United States) Flora of California Plants used in traditional Native American medicine Flora without expected TNC conservation status