''Davilla'' is a genus of

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

s belonging to the family

Dilleniaceae Dilleniaceae is a family of flowering plants with 11 genera and about 430 known species. Such a family has been universally recognized by taxonomists. It is known to gardeners for the genus ''Hibbertia'', which contains many commercially valuabl ...

. It has around 30 neotropical, species and is one of most diverse genera of

liana A liana is a long- stemmed, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the canopy in search of direct sunlight. The word ''liana'' does not refer to a ta ...

s, vines, erect or scandent (climbing) shrubs.

Description

''Davilla'' plants are classed as

lianas A liana is a long- stemmed, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the canopy in search of direct sunlight. The word ''liana'' does not refer to a ta ...

or shrubs, and they are similar in form to that of species in ''

Tetracera ''Tetracera'' is a genus of flowering plants of the Dilleniaceae family native to the tropics. Several species are lianas. Species '' Plants of the World Online'' currently includes: # '' Tetracera affinis'' Hutch. # '' Tetracera akara'' (Bu ...

'' or ''

Dillenia ''Dillenia'' is a genus of flowering plants in the family Dilleniaceae, native to tropical and subtropical regions of southern Asia, Australasia, and the Indian Ocean islands. The genus is named after the German botanist Johann Jacob Dillenius, ...

'' genus.George Don Although they can identified from other Dilleniaceae genera plants due to several features; having sepals unequal in size, with the two inner ones larger, becoming crustaceous (having a hard shell) and covering the fruit completely, a

paniculate A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are of ...

inflorescence and the fruit being a capsule. They are

hermaphroditic In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have separ ...

plants (or bisexual - bearing both male and female reproductive organs). It has leaves which are often scabrous (rough to the touch) and pubescent with simple trichomes (hairs or bristles). The petioles (leaf stalks) are winged to narrowly recurve-winged. The inflorescences panicles are terminal (at the end of branches) or axillary (at leaf junctions) in the upper nodes. The flower has 5

sepals A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...

, which are uneven. The 3 outer ones are small and 2 inners ones are larger. It has 3–6

petals Petals are modified leaves that surround the reproductive parts of flowers. They are often brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corolla''. Petals are usually ...

, which are deciduous. It has numerous

stamens The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...

and 1-2

carpels Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...

(female reproductive organ), which are capsular and contain 1 compartment. It has 2

ovules In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the fe ...

and erect and basal clavate (club-shaped) styles. It also has peltate (shield-like) and

emarginate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...

(notched at the apex) stigmas. The sepals later become leathery and begin enveloping the fruit, and simulating a globose shaped capsule. Inside the capsule, are 1-2 smooth seeds which are surrounded by an

aril An aril (pronounced ), also called an arillus, is a specialized outgrowth from a seed that partly or completely covers the seed. An arillode or false aril is sometimes distinguished: whereas an aril grows from the attachment point of the see ...

(a membranous or fleshy appendage). Not much of the

reproductive The reproductive system of an organism, also known as the genital system, is the biological system made up of all the anatomical organs involved in sexual reproduction. Many non-living substances such as fluids, hormones, and pheromones are als ...

biology of the genus Davilla is known. Noting floral visitors, among reports concerning the genus ''Davilla'', Ducke (1902) reported that bee species, ''

Halictus The genus ''Halictus'' is a large assemblage of bee species in the family Halictidae. The genus is divided into 15 subgenera, some of dubious monophyly, containing over 200 species, primarily in the Northern Hemisphere (a few species occur in So ...

'' was visiting flowers of ''Davilla rugosa'' . For the same species, Kuhlmann & Kühn (1947) indicated bees and other insects were listed as

pollinators A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the maj ...

. Croat (1978), then verified flowers of ''Davilla nitida'' were being visited by (stingless bee species) Partamona cupira (Smith 1863).

Taxonomy

The genus name of ''Davilla'' is either named in honour of Pedro Franco Dávila (1711–1786), a Peruvian and Spanish naturalist and collector, or according to

George Don George Don (29 April 1798 – 25 February 1856) was a Scottish botanist and plant collector. Life and career George Don was born at Doo Hillock, Forfar, Angus, Scotland on 29 April 1798 to Caroline Clementina Stuart and George Don (b.1756), p ...

in 1831, ''Henry Catherine Davilla'' (

Enrico Caterino Davila Enrico Caterino Davila (30 October 157626 May 1631) was an Italian historian and diplomat. Life Born in Piove di Sacco, near Padua, he was descended from a Spanish noble family. He was the youngest son of Antonio Davila, Grand Constable of Cyprus ...

) an Italian historian who died in 1599. The genus was first published and described by

Domenico Vandelli Domenico Agostino Vandelli (Padua, 8 July 1735 – Lisbon, 27 June 1816) was an Italian naturalist, who did most of his scientific work in Portugal. He studied at the University of Padua, from which he received a doctorate in Natural Philos ...

in Fl. Lusit. Brasil. Spec. collation 35 in 1788. It was then reprinted in Script. Pl. Hispan. (edited by J.J. Roemer), Vol.115 in 1796. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Davilla rugosa'' . The taxonomy of the genus ''Davilla'' was revised in 2012.

Species known

According to Kew; The genus is accepted by

United States Department of Agriculture The United States Department of Agriculture (USDA) is the United States federal executive departments, federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, ...

and the

Agricultural Research Service The Agricultural Research Service (ARS) is the principal in-house research agency of the United States Department of Agriculture (USDA). ARS is one of four agencies in USDA's Research, Education and Economics mission area. ARS is charged with ext ...

, they accept 2 species; ''Davilla nitida'' (Vahl) Kubitzki and ''Davilla rugosa'' Poir.

Distribution and habitat

Its native range is between Mexico and tropical America. It is found in the countries of;

Belize Belize (; bzj, Bileez) is a Caribbean and Central American country on the northeastern coast of Central America. It is bordered by Mexico to the north, the Caribbean Sea to the east, and Guatemala to the west and south. It also shares a wate ...

Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...

, Brazil,

Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...

Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...

Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...

Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...

French Guiana French Guiana ( or ; french: link=no, Guyane ; gcr, label=French Guianese Creole, Lagwiyann ) is an overseas departments and regions of France, overseas department/region and single territorial collectivity of France on the northern Atlantic ...

Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by H ...

Guyana Guyana ( or ), officially the Cooperative Republic of Guyana, is a country on the northern mainland of South America. Guyana is an indigenous word which means "Land of Many Waters". The capital city is Georgetown. Guyana is bordered by the ...

Honduras Honduras, officially the Republic of Honduras, is a country in Central America. The republic of Honduras is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Oce ...

Jamaica Jamaica (; ) is an island country situated in the Caribbean Sea. Spanning in area, it is the third-largest island of the Greater Antilles and the Caribbean (after Cuba and Hispaniola). Jamaica lies about south of Cuba, and west of His ...

, Mexico,

Nicaragua Nicaragua (; ), officially the Republic of Nicaragua (), is the largest country in Central America, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west. Managua is the cou ...

Panamá Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cost ...

Paraguay Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to th ...

Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...

Suriname Suriname (; srn, Sranankondre or ), officially the Republic of Suriname ( nl, Republiek Suriname , srn, Ripolik fu Sranan), is a country on the northeastern Atlantic coast of South America. It is bordered by the Atlantic Ocean to the north ...

Trinidad-Tobago Trinidad and Tobago (, ), officially the Republic of Trinidad and Tobago, is the southernmost island country in the Caribbean. Consisting of the main islands Trinidad and Tobago, and numerous much List of islands of Trinidad and Tobago, small ...

and

Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...

. The greatest ''Davilla'' species diversity is located in Brazil, up to 12 species can be found in the

Atlantic Forest The Atlantic Forest ( pt, Mata Atlântica) is a South American forest that extends along the Atlantic coast of Brazil from Rio Grande do Norte state in the northeast to Rio Grande do Sul state in the south and inland as far as Paraguay and th ...

of Brazil.

Habitat

It is found in moist or wet forests, often in hilly pine forests. Also in forest clearings and borders. It grows at altitudes of above sea level.

Uses

''Davilla elliptica'' and ''Davilla nitida'' as well as ''

Alchornea glandulosa ''Alchornea glandulosa'' is a tree species of the Acalyphoideae native to South America, growing in southern Brazil from Minas Gerais to Rio Grande do Sul. It is locally known as ''tamanqueiro'', ''tapiá'' or ''amor seco''. This gnarled tree gro ...

'' (a tree species), have properties that could be used in the treatment of

peptic ulcers Peptic ulcer disease (PUD) is a break in the inner lining of the stomach, the first part of the small intestine, or sometimes the lower esophagus. An ulcer in the stomach is called a gastric ulcer, while one in the first part of the intestines i ...

. The leaves of ''Davilla elliptica'' have been used in

folk medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...

to treat diseases such as inflammation and other ulcers.

Threats

''Davilla glaziovii'' is included on the

red list The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biologi ...

of Brazil, due to habitat loss and predatory

extractivism Extractivism is the process of extracting natural resources from the Earth to sell on the world market. It exists in an economy that depends primarily on the extraction or removal of natural resources that are considered valuable for exportation w ...

(the extracting natural resources).

References

{{Taxonbar, from=Q5241770 Dilleniaceae Eudicot genera Plants described in 1796 Flora of South America