''Dasygrammitis'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...

s in the family

Polypodiaceae Polypodiaceae is a family of ferns. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family includes around 65 genera and an estimated 1,650 species and is placed in the order Polypodiales, suborder Polypodiineae. A broader ...

, subfamily

Grammitidoideae Grammitidoideae is a subfamily of the fern family Polypodiaceae, whose members are informally known as grammitids. It comprises a clade of about 750 species. They are distributed over higher elevations in both the Old and New World. This group w ...

, according to the

Pteridophyte Phylogeny Group The Pteridophyte Phylogeny Group, or PPG, is an informal international group of systematic botanists who collaborate to establish a consensus on the classification of pteridophytes (lycophytes and ferns) that reflects knowledge about plant relation ...

classification of 2016 (PPG I). It is known from

Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...

through southeast Asia and the Philippines to Polynesia.

Description

The rhizomes are radially symmetric (without distinct upper and lower surfaces) and bear whorls of stipes, which lack a joint at the point of attachment. The rhizome scales are red-brown, of uniform color, and usually glossy. They have unbranched, red-brown hairs on their edges. Hairs, where present, are unbranched and branched, and brown in color. The leaf blades are usually

pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...

in cutting, rarely

pinnatifid Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common Anatomical terms of location#Axes, axis. Pinnation occurs in biological morphology (biology), morphology, in Crysta ...

or bipinnatifid, bearing free veins which lack

hydathode A hydathode is a type of pore, commonly found in angiosperms, that secretes water through pores in the epidermis or leaf margin, typically at the tip of a marginal tooth or serration. Hydathodes occur in the leaves of submerged aquatic plants s ...

s at their terminus. Sori are borne in two rows beneath pinnae or lobes. The sori are circular to elliptic in shape; the

sporangia A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point in their life cy ...

either lack hairs or have one to two red-brown hairs.

Taxonomy

The genus was first described by Barbara Parris in 2007 to receive some of the species of the genus ''

Grammitis ''Grammitis'' (dwarf polypody) is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It had formerly been placed in the family Grammitidace ...

'', which as then circumscribed was artificial and contained a number of disparate lineages. The name "''Dasygrammitis''" means "shaggy ''Grammitis''", referring to the shaggy hairs often present on the frond in this group of former ''Grammitis'' species. Parris initially placed six species in the genus. A phylogenetic study of the grammitids found that ''Dasygrammitis'' was sister to a clade consisting of ''

Calymmodon ''Calymmodon'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Its known range is Sri Lanka, Thailand, Vietnam, Taiwan, Malesia, Austra ...

'', ''

Micropolypodium ''Micropolypodium'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is native to China, Japan, and the Philippines. Species Neotro ...

'', '' Scleroglossum'', '' Tomophyllum'', and '' Xiphopterella''. These six genera share a radially symmetric rhizome, which may be a

synapomorphy In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have ...

for this clade (although the trait appears in other grammitid taxa, probably independently).

Species

, the ''Checklist of Ferns and Lycophytes of the World'' accepted the following species: *'' Dasygrammitis brevivenosa'' (Alderw.) Parris *'' Dasygrammitis crassifrons'' (Baker) Parris *'' Dasygrammitis fuscata'' (Blume) Parris *'' Dasygrammitis kinabaluensis'' Parris *''

Dasygrammitis malaccana ''Dasygrammitis'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Sri Lanka through southeast Asia and the Philippines ...

'' (Baker) Parris *''

Dasygrammitis mollicoma ''Dasygrammitis'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Sri Lanka through southeast Asia and the Philippines ...

'' (Nees & Blume) Parris *''

Dasygrammitis purpurascens ''Dasygrammitis'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Sri Lanka through southeast Asia and the Philippines ...

'' (Nadeaud) Parris *'' Dasygrammitis seramensis'' Parris

References

{{Taxonbar, from=Q17197627 Polypodiaceae Fern genera