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''Dares ulula'' is a species of
stick insects The Phasmatodea (also known as Phasmida, Phasmatoptera or Spectra) are an order of insects whose members are variously known as stick insects, stick-bugs, walking sticks, stick animals, or bug sticks. They are also occasionally referred to as ...
. Like most other members of the genus '' Dares'', the species is native to Borneo, more precisely in the northwest of the island. The males are extremely prickly even for the representatives of the genus '' Orestes''. Females are colored with a relatively high contrast.


Characteristics

The species shows roughly the typical habitus for the genus. It is one of the largest representatives of the genus ''Dares''. Females grow to be long. They are more contrasting than the females of other species. In addition to areas with different, mostly dark brown tones, there are also areas that are yellowish, orange-brown and black-brown in color. These areas can form wide transverse bands on the abdomen. The tubercles that are on the body surface are more distinct and higher in them than in many other species. Sometimes they are so pronounced that they form small, flap-like
lobes Lobe may refer to: People with the name * Lobe (surname) Science and healthcare * Lobe (anatomy) * Lobe, a large-scale structure of a radio galaxy * Glacial lobe, a lobe-shaped glacier * Lobation, a characteristic of the nucleus of certain ...
or even short, blunt spines. The antennae consist of 25 segments. Their basal segment (scapus) is flat and have clear teeth on the outside. The abdomen is widest in the middle. The males remain smaller than the females at in length, but are also very contrasting in color. While the head and legs are medium brown, the coloration of the thorax and the abdomen is dominated by dark brown and yellow to orange longitudinal stripes. The spines on the head and thorax correspond in their arrangement to that of ''
Dares verrucosus ''Dares verrucosus'' is a species of Phasmatodea, stick insects. Like most other members of the genus ''Dares (insect), Dares'', the species is native to Borneo, more precisely in the north of the island. Characteristics Both sexes have the t ...
'', but are usually much longer than this. The males have four spines on their heads, at the front edge of the mesonotum and in the rear area of the meso- and metanotum there are a total of three further pairs of spines. To the side of it there is another spine on the pleura. There are no spines on the abdomen. The antennae of the males consist of only 23 segments. Their basal segment is also flat and have clear and relatively pointed teeth on the outside.


Distribution area and lifestyle

''Dares ulula'' is widespread in the
Malay Malay may refer to: Languages * Malay language or Bahasa Melayu, a major Austronesian language spoken in Indonesia, Malaysia, Brunei and Singapore ** History of the Malay language, the Malay language from the 4th to the 14th century ** Indonesi ...
state of Sarawak in the northwest of the island of Borneo. Many previously known origins were destroyed by urbanization and the creation of new plantations. With their defensive behavior and nocturnal lifestyle, this species differs little from other ''Dares'' species. Their defense strategy is based on phytomimesis. When touched, the animals let themselves fall to the ground, where they play dead until the threat has passed. The females lay their eggs on the ground. The eggs are not quite spherical and almost hairless. They are approximately long and wide. Their micropylar plate has four arms and the shape of an "X". The arms reach far around the egg. The nymphs are usually very dark in color and show conspicuously bright, mostly yellow-orange patterns around the resulting spines that increase with age or their stage of development.


Taxonomy

John Obadiah Westwood described the species as ''Acanthoderus ulula'' in 1859. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''ulula'' chosen by him means "owl". The female nymph on which his description is based was collected by D. Wallace in Sarawak. It is deposited as holotype of the species in the Oxford University Museum of Natural History. In 1875 Carl Stål transferred the species as a second representative next to ''
Dares validispinus ''Dares validispinus'' is a species of stick insects. Like most other members of the genus ''Dares'', the species is native to Borneo, more precisely in the northwest of the island. Characteristics The species shows the typical habitus for t ...
'' in the genus ''Dares'' which he had newly established. Josef Redtenbacher described two species in 1906, both of which in 1935 by Klaus Günther
synonymed A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
to ''Dares ulula''. One of these synonyms is ''Dares calamita''. The females to which the description referred were deposited in the Hungarian Natural History Museum in Budapest, but are considered destroyed. The second synonym is ''Dares corticinus''. Here Redtenbacher described a very young male nymph, which is deposited as holotype at the Muséum national d'histoire naturelle in Paris.


Terraristic

The first live specimens of this species came to Europe in 1991. Both
Philip Edward Bragg Philip, also Phillip, is a male given name, derived from the Greek (''Philippos'', lit. "horse-loving" or "fond of horses"), from a compound of (''philos'', "dear", "loved", "loving") and (''hippos'', "horse"). Prominent Philips who popularize ...
and Patrick van der Stigchel and even Ian Abercrombie brought specimens from Mount Serapi that year. ''Dares ulula'' was the third ''Dares'' species that got into the European terrariums. Since then the species has been introduced several times. It is considered to be delicate to care for, but because of its appearance it is the most sought-after representative of the genus. Like the other species of the genus, it prefers a higher humidity, which can be achieved by a layer of soil covered with moist
moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hor ...
. Besides leaves of brambles and many other
Rosaceae Rosaceae (), the rose family, is a medium-sized family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus ''Rosa''. Among the most species-rich genera are ''Alchemilla'' (270), ''Sorbus ...
, the leaves of various Betulaceae are also eaten. The
Phasmid Study Group The Phasmid Study Group is a group for professional and amateur entomologists who are interested in the order Phasmatodea, i.e. stick and leaf insects, known as "phasmids", as well as other interested persons. Over 40 species In biology, a s ...
listed ''Dares ulula'' under PSG number 117.


References

Bragg, P. E. (2001). ''Phasmids of Borneo'', Natural History Publications (Borneo) Sdn. Bhd., Kota Kinabalu, Sabah, Malaysia, pp. 126–153, Dräger, H. (2012). ''Gespenstschrecken der Familie Heteropterygidae Kirby, 1896 (Phasmatodea) – ein Überblick über bisher gehaltene Arten, Teil 2: Die Unterfamilie Dataminae Rehn & Rehn, 1839'', ZAG Phoenix, Nr. 5 Juni 2012 Jahrgang 3(1), pp. 22–45, Westwood, J. O. (1859).
Catalog of the orthopterous insects in the collection of the British Museum. Part I. Phasmidae.
' London, p. 53 & plate 26 Fig. 1
Josef Redtenbacher (1906).
Die Insektenfamilie der Phasmiden. Vol. 1. Phasmidae Areolatae
'. Verlag Wilhelm Engelmann, Leipzig, pp. 53–56
phasmatodea.com
by Hennemann, F. H.; Conle, O. V.; Kneubühler, B. & Valero, P.
Bragg, P. E. (1998). ''A revision of the Heteropteryginae (Insecta: Phasmida: Bacillidae) of Borneo, with the description of a new genus and ten new species'', Zoologische Verhandelingen, Leiden 316, pp. pp. 60-64 & 115-122. ,
Download version
/ref> Brock, P. D.; Büscher, T. H. & Baker, E. W.
Species File Online
'. Version 5.0. (accessdate 1 August 2021)
Phasmid Study Group Culture List
/ref>


External links

* * {{Taxonbar, from=Q1103974 Phasmatodea Phasmatodea of Asia Insects described in 1859