Daka Skull
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The Daka calvaria, otherwise known as the Daka skull (a skull cap with the cranial base), or specimen number BOU-VP-2/66, is a '' Homo erectus'' specimen from the Daka Member of the
Bouri Formation The Bouri Formation is a sequence of sedimentary deposits that is the source of australopithecine and Homo (that is, hominin) fossils, artifacts, and bones of large mammals with cut marks from butchery with tools by early hominins. It is locat ...
in the Middle Awash Study Area of the
Ethiopian Rift Valley The Great Rift Valley of Ethiopia, (or Main Ethiopian Rift or Ethiopian Rift Valley) is a branch of the East African Rift that runs through Ethiopia in a southwest direction from the Afar Triple Junction. In the past, it was seen as part of a "Gr ...
. It is reported that the metrics and morphological attributes of the Daka calvaria center it firmly within the '' Homo erectus'' lineage. The resemblance to the Asian counterparts indicates that early African and Eurasian fossil
hominids The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); ''Gorilla'' (the ...
represent a subdivision of a widespread paleospecies. Daka's anatomical intermediacy between "earlier and later African fossils provides evidence of evolutionary change. Its temporal and geographic position indicates that African
H. erectus ''Homo erectus'' (; meaning "upright man") is an extinct species of archaic human from the Pleistocene, with its earliest occurrence about 2 million years ago. Several human species, such as '' H. heidelbergensis'' and '' H. antecessor' ...
was the ancestor of ''
Homo sapiens Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...
''.” It was discovered in 1997 by Henry Gilbert. With it are several other ''
H. erectus ''Homo erectus'' (; meaning "upright man") is an extinct species of archaic human from the Pleistocene, with its earliest occurrence about 2 million years ago. Several human species, such as '' H. heidelbergensis'' and '' H. antecessor' ...
'' specimens (leg bones, cranial fragments, and a toothless mandible), a large assortment of
Acheulean Acheulean (; also Acheulian and Mode II), from the French ''acheuléen'' after the type site of Saint-Acheul, is an archaeological industry of stone tool manufacture characterized by the distinctive oval and pear-shaped "hand axes" associated ...
industry stone tools, and several hundred animal fossils. The locality dates to about one million years old.


Origin

The one-million-year-old Homo sample BOU-VP-2/66 from the Dakanihylo Member, referred to as “Daka” calvaria, was discovered in Bouri,
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
, found in the Middle Awash region of the Afar Rift. This area has, not surprisingly, been the source of many human ancestor fossils, with ages ranging from half a million years old to more than 6 million years old. The resemblance of the Daka Skull to Asian counterparts indicates that the early Eurasian and African fossil hominids represent demes of a widespread palaeospecies. The anatomical intermediacy of Daka between earlier and later African fossils provides evolutionary change evidence. There is a consensus that there is a “lack of calvarial evidence for a deep phylo-genetic division between the African and Asian fossils.” According to researchers, a phenetic approach, quantifying overall similarity of single specimens, found that the Daka calvaria shares affinities with two fossils from the
Koobi Fora Koobi Fora refers primarily to a region around Koobi Fora Ridge, located on the eastern shore of Lake Turkana in the territory of the nomadic Gabbra people. According to the National Museums of Kenya, the name comes from the Gabbra language: ...
region of Africa,
KNM-ER 3733 KNM ER 3733 is a fossilized hominid cranium of the extinct hominid ''Homo ergaster'', alternatively referred to as African ''Homo erectus''. It was discovered in 1975 in Koobi Fora, Kenya, right next to Lake Turkana, in a survey led by Richard L ...
and KNM-ER 3883. These are attributed to ''
Homo ergaster ''Homo ergaster'' is an extinct species or subspecies of archaic humans who lived in Africa in the Early Pleistocene. Whether ''H. ergaster'' constitutes a species of its own or should be subsumed into '' H. erectus'' is an ongoing and unresol ...
'', which is very different from '' Homo erectus''.


Cranial morphology

The Daka Calvaria has a cranial capacity of 995cc (for reference, a chimp has about 375cc and a human about 1,200cc). Through the use of computed tomographic (CT) imagery of the Daka calvaria, observations have been made on the distortion, subcortical structures and endocranial features of this skull fossil. According to the CT-derived metrics for the Daka calvaria, the occipital Asterion thickness was 10.6 mm, with the temporal Asterion thickness measuring at 10.2mm. The Calvarial height above g-l, above g-op and above n-op, were measured at 51mm, 73mm and 80mm, respectively; Endocranial capacity was found to be around 986ml. The strong flexion that is apparent in Daka strengthens the theory that midsagittal cranial base flexion was strong in '' Homo erectus'' in general. Due to the age and condition of the Daka skull, the fossil is not fully intact. The right side of the calvaria is slightly more damaged and there is a relatively large space in the posterior region of the mastoid area near the Asterion. Researchers found that the mastoid process of the artifact are both damaged extensively. “Neither side preserves the inferior portions, but pneumatization of the temporal bone superior to the mastoid tips extends superiorly nearly to Asterion on both sides.” The temporals on the skull are not as inflated as those of apes, with smaller individual cells. The Daka calvaria temporal squamae measure at approximately 5.7mm thick centrally. The posterior temporal is approximately 10.6mm thick adjacent to the occipitomastoid suture.


References


External links


Daka Calvarium
- UC Press Journal Article {{Commons, Homo erectus skulls Homo erectus fossils Individual human heads, skulls and brains 1997 archaeological discoveries Archaeological discoveries in Ethiopia