''Cyrtopleura costata'', or the angel wing clam, is a

bivalve Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...

mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is esti ...

in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Pholadidae Pholadidae, known as piddocks or angelwings, are a family of bivalve molluscs similar to a clam. Background Piddocks are unique in that each side of their shells is divided into 2 or 3 separate sections. Furthermore, one of the piddock's shells ...

. It is found in shallow parts of the northwest Atlantic and also in the North Sea of Scotland coastline and west coast of the Adriatic Sea by a remote area in the Marche region in central Italy, living in the seabed, where it digs its burrows on a very slow revolving movement for years through soft sand and mud always to a max depth of 8ft but always below at the lowest tide.''Cyrtopleura costata''
Smithsonian Marine Station. Retrieved 2011-11-30.

Description

''Cyrtopleura costata'' has a pair of brittle, asymmetric white valves and can grow to about in length. The anterior end is elongated and has a rounded point which is used for digging through the substrate. The posterior end is truncated and rounded and near the beak has an apophysis, a wing-like flange, which helps provide an attachment for the foot muscles. On the anterior side of the beak, the margin is smooth and bent slightly upwards. The whole valve has finely sculptured radial ribs which intercept with a series of concentric growth rings parallel with the margin. In the living animal, the valves are covered by the

periostracum The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods and ...

, a thin grey protective layer of protein which is part of the shell. This layer has usually been stripped away by sand and surf by the time that the empty shells are washed up on the beach, and the valves are usually found singly, because the muscles that hold them together are weak. Internally, the live mollusc has a powerful muscular foot and a pair of long, fused siphons. These siphons are unable to retract back into the shell and as a result, the two valves permanently gape apart.Angel Wings: Look for the “Spoon”
Retrieved 2011-11-30.

Distribution

''Cyrtopleura costata'' is found in shallow seas in the north east

Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...

between

Cape Cod Cape Cod is a peninsula extending into the Atlantic Ocean from the southeastern corner of mainland Massachusetts, in the northeastern United States. Its historic, maritime character and ample beaches attract heavy tourism during the summer mont ...

and the

Gulf of Mexico The Gulf of Mexico ( es, Golfo de México) is an oceanic basin, ocean basin and a marginal sea of the Atlantic Ocean, largely surrounded by the North American continent. It is bounded on the northeast, north and northwest by the Gulf Coast of ...

. It is also found in the

West Indies The West Indies is a subregion of North America, surrounded by the North Atlantic Ocean and the Caribbean Sea that includes 13 independent island countries and 18 dependencies and other territories in three major archipelagos: the Greater A ...

Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...

and as far south as

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

. It is commonest in the

intertidal zone The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species o ...

and just below low water mark.

Biology

''Cyrtopleura costata'' lives beneath the surface of the sea bed. It is able to bore through sand, mud, wood, clay and even soft rock using a twisting motion of its pointed, anterior end assisted by jets of water ejected from the mantle cavity. It is a

filter feeder Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedin ...

. The siphons extend to the surface of the substrate and water is drawn in through one and expelled through the other with

microalgae Microalgae or microphytes are microscopic algae invisible to the naked eye. They are phytoplankton typically found in freshwater and marine systems, living in both the water column and sediment. They are unicellular species which exist indiv ...

and

zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...

being filtered out as the water passes through the gills. Respiration takes place at the same time. Cilia then waft the food particles to the mouth. Spawning usually takes place in summer.

Gamete A gamete (; , ultimately ) is a haploid cell that fuses with another haploid cell during fertilization in organisms that reproduce sexually. Gametes are an organism's reproductive cells, also referred to as sex cells. In species that produce t ...

s are passed out of the exhalent siphon and fertilisation takes place externally. After hatching,

eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...

develop into

veliger A veliger is the planktonic larva of many kinds of sea snails and freshwater snails, as well as most bivalve molluscs (clams) and tusk shells. Description The veliger is the characteristic larva of the gastropod, bivalve and scaphopod ...

larvae which are

plankton Plankton are the diverse collection of organisms found in Hydrosphere, water (or atmosphere, air) that are unable to propel themselves against a Ocean current, current (or wind). The individual organisms constituting plankton are called plankt ...

ic. After 16 to 21 days these undergo

metamorphosis Metamorphosis is a biological process by which an animal physically develops including birth or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and differentiation. Some inse ...

into a pediveliger stage and settle out onto a soft substrate such as sand. There they become juveniles and start burrowing.

Uses

''Cyrtopleura costata'' is used as food in

Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...

and

Puerto Rico Puerto Rico (; abbreviated PR; tnq, Boriken, ''Borinquen''), officially the Commonwealth of Puerto Rico ( es, link=yes, Estado Libre Asociado de Puerto Rico, lit=Free Associated State of Puerto Rico), is a Caribbean island and Unincorporated ...

. It is a fast-growing species, and it has been investigated for possible use in commercial

aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. lot ...

. Under optimal conditions it was found that the larvae were ready to undergo

after 12 days, and settlement could be triggered by use of a dilute solution of

epinephrine Adrenaline, also known as epinephrine, is a hormone and medication which is involved in regulating visceral functions (e.g., respiration). It appears as a white microcrystalline granule. Adrenaline is normally produced by the adrenal glands and ...

. The clams could reach a marketable size of within 6 months.

References

{{Taxonbar, from=Q3014578 Pholadidae Molluscs described in 1758 Taxa named by Carl Linnaeus