Coptoclavidae

Coptoclavidae is an extinct family of aquatic beetles in the suborder Adephaga. The Coptoclavidae lived from the Late Triassic to the Early Cretaceous. Coptoclavidae is a member of the adephagan clade Dytiscoidea, which contains other aquatic beetles. Suggested reasons for their extinction to include the rise of teleost fish, or competition with Gyrinidae and Dytiscidae, which possess defensive secretions and sucking channels in the mandibles of larvae, which coptoclavids likely lacked. It has been suggested that the genus '' Timarchopsis'' and the subfamily Timarchopsinae are only distantly related to other coptoclavids based on cladistic analysis, with ''Timarchopsis'' being more closely related to geadephagans like carabids and trachypachids instead. Another study also suggested similarly for '' Coptoclavisca'' and possibly other ''c''optoclaviscines.

Taxonomy

* †'' Agrascapha'' Lin 1992 Huangshanjie Formation, China, Carnian
* †'' Amblycephalonius'' Bode 1953 Posidonia Shale, Germany, Toarcian
* †subfamily Charonoscaphinae Ponomarenko 1977
** †'' Charonoscapha'' Ponomarenko 1977 Karabastau Formation, Kazakhstan, Callovian
** †'' Charonoscaphidia'' Ponomarenko 1977 Karabastau Formation, Kazakhstan, Callovian
* †'' Coptoclavia'' Ping 1928 Cassange Group, Upper Triassic
* †subfamily Coptoclavinae Ponomarenko 1961
** †'' Bolbonectes'' Ponomarenko 1987
*** †''Bolbonectes intermedius'' Ponomarenko 1987 Byankino Formation, Russia, Tithonian Leskovskaya Formation, Russia, Barremian
*** †''Bolbonectes occidentalis'' Ponomarenko 1993 Ichetuy Formation, Russia, Oxfordian
*** †''Bolbonectus lithographicus'' Ponomarenko and Martínez-Delclòs 2000 La Pedrera de Rúbies Formation, Spain, Barremian
** †'' Coptoclava'' Ping 1928 Ustkarskaya Formation, Russia, Hauterivian Dabeigou Formation