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The
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideall ...
Asteraceae, alternatively Compositae, consists of over 32,000 known species of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s in over 1,900 genera within the order
Asterales Asterales () is an order of dicotyledonous flowering plants that includes the large family Asteraceae (or Compositae) known for composite flowers made of florets, and ten families related to the Asteraceae. While asterids in general are char ...
. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae were first described in the year 1740. The number of species in Asteraceae is rivaled only by the
Orchidaceae Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Along with the Asteraceae, they are one of the two largest families of flowering ...
, and which is the larger family is unclear as the quantity of
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extin ...
species in each family is unknown. Most species of Asteraceae are
annual Annual may refer to: *Annual publication, periodical publications appearing regularly once per year **Yearbook **Literary annual *Annual plant *Annual report *Annual giving *Annual, Morocco, a settlement in northeastern Morocco *Annuals (band), a ...
,
biennial Biennial means (an event) lasting for two years or occurring every two years. The related term biennium is used in reference to a period of two years. In particular, it can refer to: * Biennial plant, a plant which blooms in its second year and th ...
, or
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
herbaceous plants, but there are also shrubs, vines, and trees. The family has a widespread distribution, from subpolar to tropical regions in a wide variety of habitats. Most occur in hot desert and cold or hot semi-desert climates, and they are found on every continent but
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
. The primary common characteristic is the existence of sometimes hundreds of tiny individual
florets This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
which are held together by protective
involucre In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or o ...
s in
flower head A pseudanthium (Greek for "false flower"; ) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, compos ...
s, or more technically, ''capitula''. The oldest known fossils are pollen grains from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
(
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian ...
to
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
) of Antarctica, dated to
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
(mya). It is estimated that the
crown group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. ...
of Asteraceae evolved at least 85.9 mya (Late Cretaceous,
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. T ...
) with a stem node age of 88–89 mya (Late Cretaceous,
Coniacian The Coniacian is an age or stage in the geologic timescale. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series and spans the time between 89.8 ± 1 Ma and 86.3 ± 0.7 Ma (million years ago). The Coniacian is preceded by t ...
). Asteraceae is an economically important family, providing food staples, garden plants, and herbal medicines. Species outside of their native ranges can be considered weedy or invasive.


Description

Members of the Asteraceae are mostly herbaceous plants, but some shrubs, vines, and trees (such as ''
Lachanodes arborea ''Lachanodes arborea'', the she cabbage tree, is a small tree in the family Asteraceae. It is an endangered endemic of the island of Saint Helena in the South Atlantic Ocean. It is now extinct in the wild. See also *Flora of St Helena The flora ...
'') do exist. Asteraceae species are generally easy to distinguish from other plants because of their unique
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed on ...
and other shared characteristics, such as the joined anthers of the
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s. However, determining genera and species of some groups such as ''
Hieracium ''Hieracium'' (), known by the common name hawkweed and classically as (from ancient Greek ιεράξ, 'hawk'), is a genus of flowering plant in the family Asteraceae, and closely related to dandelion (''Taraxacum''), chicory ('' Cichorium'' ...
'' is notoriously difficult (see " damned yellow composite" for example).


Roots

Members of the family Asteraceae generally produce
taproot A taproot is a large, central, and dominant root from which other roots sprout laterally. Typically a taproot is somewhat straight and very thick, is tapering in shape, and grows directly downward. In some plants, such as the carrot, the taproo ...
s, but sometimes they possess fibrous root systems. Some species have underground stems in the form of caudices or
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow ho ...
s. These can be fleshy or woody depending on the species.


Stems

Stems are herbaceous, aerial, branched, and cylindrical with glandular hairs, generally erect, but can be prostrate to ascending. The stems can contain secretory canals with
resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on natu ...
, or
latex Latex is an emulsion (stable dispersion) of polymer microparticles in water. Latexes are found in nature, but synthetic latexes are common as well. In nature, latex is found as a milky fluid found in 10% of all flowering plants (angiosperms ...
which is particularly common among the
Cichorioideae The Cichorioideae are a subfamily of the family Asteraceae of flowering plants. Familiar members of Cichorioideae include lettuce, dandelions, chicory and '' Gazania'' species. The subfamily comprises about 240 genera and about 2900 species. It i ...
.


Leaves

Leaves can be
alternate Alternative or alternate may refer to: Arts, entertainment and media * Alternative (''Kamen Rider''), a character in the Japanese TV series ''Kamen Rider Ryuki'' * ''The Alternative'' (film), a 1978 Australian television film * ''The Alternative ...
, opposite, or whorled. They may be simple, but are often deeply lobed or otherwise incised, often conduplicate or revolute. The margins also can be
entire Entire may refer to: * Entire function, a function that is holomorphic on the whole complex plane * Entire (animal) Neutering, from the Latin ''neuter'' ('of neither sex'), is the removal of an animal's reproductive organ, either all of it or a c ...
or toothed. Resin or latex also can be present in the leaves.


Inflorescences

Nearly all Asteraceae bear their flowers in dense flower heads called ''capitula''. They are surrounded by involucral bracts, and when viewed from a distance, each capitulum may appear to be a single flower. Enlarged outer (peripheral) flowers in the capitulum may resemble petals, and the involucral bracts may look like a calyx.


Floral heads

In plants of the family Asteraceae, what appears to be a single flower is actually a cluster of much smaller flowers. The overall appearance of the cluster, as a single flower, functions in attracting
pollinators A pollinator is an animal that moves pollen from the male anther of a flower to the female carpel, stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are ...
in the same way as the structure of an individual flower in some other plant families. The older family name, Compositae, comes from the fact that what appears to be a single flower is actually a ''composite'' of smaller flowers. The "petals" or "sunrays" in a sunflower head are actually individual strap-shaped flowers called ''ray flowers'' or ''ray florets'', and the "sun disk" is made of smaller circular shaped individual flowers called ''disc flowers'' or ''disk florets''. The word ''aster'' means "star" in Greek, referring to the appearance of some family members, as a star surrounded by rays. The cluster of flowers that may appear to be a single flower is called a ''head''. The entire head may move tracking the sun, like a "smart" solar panel, which maximizes reflectivity of the whole unit and can thereby attract more pollinators. On the outside the flower heads are small bracts that look like scales. These are called ''
phyllaries In botanical terminology, a phyllary, also known an involucral bract or tegule, is a single bract of the involucre of a composite flower. The involucre is the grouping of bracts together. Phyllaries are reduced leaf-like structures that form one o ...
'', and together they form the
involucre In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or o ...
that protects the individual flowers in the head before they open. The individual heads have the smaller individual flowers (florets) arranged on a round or dome-like structure called the ''receptacle''. The florets mature first at the outside, moving toward the center, with the youngest in the middle. The individual flowers in a head have 5 fused
petal Petals are modified leaves that surround the reproductive parts of flowers. They are often brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corolla''. Petals are usually ...
s (rarely 4), but instead of
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined ...
s, they have threadlike, hairy, or bristly structures singularly called a '' pappus'', plural ''pappi'', which surround the fruit and can stick to animal fur or be lifted by wind, aiding in seed dispersal. The whitish fluffy head of a dandelion, commonly blown on by children, is made of pappi with tiny seeds attached at the ends. The pappi provide a parachute like structure to help the seed be carried away in the wind. A ''ray flower'' is a 3-tipped (3-lobed), strap-shaped, individual flower in the head of some members of the family Asteraceae. Sometimes a ray flower is 2-tipped (2-lobed). The corolla of the ray flower may have 2 tiny teeth opposite the 3-lobed strap, or tongue, indicating evolution by fusion from an originally 5-part corolla. Sometimes, the 3:2 arrangement is reversed, with 2 tips on the tongue, and 0 or 3 tiny teeth opposite the tongue. A ''ligulate flower'' is a 5-tipped, strap-shaped, individual flower in the heads of other members. A ''ligule'' is the strap-shaped tongue of the corolla of either a ray flower or of a ligulate flower. A ''disk flower'' (or ''disc flower'') is a radially symmetric (i.e., with identical shaped petals arranged in circle around the center) individual flower in the head, which is ringed by ray flowers when both are present. Sometimes ray flowers may be slightly off from radial symmetry, or weakly bilaterally symmetric, as in the case of desert pincushions '' Chaenactis fremontii''. A ''radiate head'' has disc flowers surrounded by ray flowers. A ''ligulate head'' has all ligulate flowers. When a sunflower family flower head has only disc flowers that are sterile, male, or have both male and female parts, it is a ''discoid head''. ''Disciform heads'' have only disc flowers, but may have two kinds (male flowers and female flowers) in one head, or may have different heads of two kinds (all male, or all female). ''Pistillate heads'' have all female flowers. ''Staminate heads'' have all male flowers. Sometimes, but rarely, the head contains only a single flower, or has a single flowered pistillate (female) head, and a multi-flowered male staminate (male) head.


Floral structures

The distinguishing characteristic of Asteraceae is their
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed on ...
, a type of specialised, composite flower head or ''
pseudanthium A pseudanthium (Greek for "false flower"; ) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, compos ...
'', technically called a calathium or ''
capitulum capitulum (plural capitula) may refer to: *the Latin word for chapter ** an index or list of chapters at the head of a gospel manuscript ** a short reading in the Liturgy of the Hours *** derived from which, it is the Latin for the assembly known ...
'', that may look superficially like a single flower. The ''capitulum'' is a contracted
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
composed of numerous individual
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s, called ''florets'', all sharing the same receptacle. A set of bracts forms an
involucre In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or o ...
surrounding the base of the capitulum. These are called "phyllaries", or "involucral bracts". They may simulate the sepals of the pseudanthium. These are mostly herbaceous but can also be brightly coloured (e.g. ''
Helichrysum The genus ''Helichrysum'' consists of an estimated 600 species of flowering plants in the sunflower family (Asteraceae). The type species is ''Helichrysum orientale''. They often go by the names everlasting, immortelle, and strawflower. The name ...
'') or have a scarious (dry and membranous) texture. The phyllaries can be free or fused, and arranged in one to many rows, overlapping like the tiles of a roof (''imbricate'') or not (this variation is important in identification of tribes and genera). Each floret may be subtended by a bract, called a "palea" or "receptacular bract". These bracts are often called " chaff". The presence or absence of these bracts, their distribution on the receptacle, and their size and shape are all important diagnostic characteristics for genera and tribes. The florets have five petals fused at the base to form a corolla tube and they may be either
actinomorphic Floral symmetry describes whether, and how, a flower, in particular its perianth, can be divided into two or more identical or mirror-image parts. Uncommonly, flowers may have no axis of symmetry at all, typically because their parts are spirall ...
or
zygomorphic Floral symmetry describes whether, and how, a flower, in particular its perianth, can be divided into two or more identical or mirror-image parts. Uncommonly, flowers may have no axis of symmetry at all, typically because their parts are spirall ...
. ''Disc florets'' are usually actinomorphic, with five petal lips on the rim of the corolla tube. The petal lips may be either very short, or long, in which case they form deeply lobed petals. The latter is the only kind of floret in the
Carduoideae Carduoideae is the thistle subfamily of the Asteraceae, or sunflower family, of flowering plants. It comprises a number of tribes in various circumscriptions of the family, in addition to the Cynareae. Takhtajan, according to Reveal, includes ...
, while the first kind is more widespread. ''Ray florets'' are always highly zygomorphic and are characterised by the presence of a ''ligule'', a strap-shaped structure on the edge of the corolla tube consisting of fused petals. In the Asteroideae and other minor subfamilies these are usually borne only on florets at the circumference of the capitulum and have a 3+2 scheme – above the fused corolla tube, three very long fused petals form the ligule, with the other two petals being inconspicuously small. The Cichorioideae has only ray florets, with a 5+0 scheme – all five petals form the ligule. A 4+1 scheme is found in the Barnadesioideae. The tip of the ligule is often divided into teeth, each one representing a petal. Some marginal florets may have no petals at all (filiform floret). The calyx of the florets may be absent, but when present is always modified into a pappus of two or more teeth, scales or bristles and this is often involved in the dispersion of the seeds. As with the bracts, the nature of the pappus is an important diagnostic feature. There are usually four or five
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s. The filaments are fused to the corolla, while the anthers are generally connate (''syngenesious'' anthers), thus forming a sort of tube around the style (''theca''). They commonly have basal and/or apical appendages. Pollen is released inside the tube and is collected around the growing style, and then, as the style elongates, is pushed out of the tube (''nüdelspritze''). The
pistil Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pistils ...
consists of two connate
carpel Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils' ...
s. The
style Style is a manner of doing or presenting things and may refer to: * Architectural style, the features that make a building or structure historically identifiable * Design, the process of creating something * Fashion, a prevailing mode of clothing ...
has two lobes. Stigmatic tissue may be located in the interior surface or form two lateral lines. The
ovary The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. ...
is inferior and has only one
ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the fem ...
, with basal
placentation Placentation refers to the formation, type and structure, or arrangement of the placenta. The function of placentation is to transfer nutrients, respiratory gases, and water from maternal tissue to a growing embryo, and in some instances to remove ...
.


Fruits and seeds

In members of the Asteraceae the fruit is
achene An achene (; ), also sometimes called akene and occasionally achenium or achenocarp, is a type of simple dry fruit produced by many species of flowering plants. Achenes are monocarpellate (formed from one carpel) and indehiscent (they do not ope ...
-like, and is called a ''cypsela'' (plural ''cypselae''). Although there are two fused carpels, there is only one locule, and only one
seed A seed is an Plant embryogenesis, embryonic plant enclosed in a testa (botany), protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, includ ...
per fruit is formed. It may sometimes be winged or spiny because the pappus, which is derived from calyx tissue often remains on the fruit (for example in
dandelion ''Taraxacum'' () is a large genus of flowering plants in the family Asteraceae, which consists of species commonly known as dandelions. The scientific and hobby study of the genus is known as taraxacology. The genus is native to Eurasia and Nor ...
). In some species, however, the pappus falls off (for example in ''
Helianthus ''Helianthus'' () is a genus comprising about 70 species of annual and perennial flowering plants in the daisy family Asteraceae commonly known as sunflowers. Except for three South American species, the species of ''Helianthus'' are native to N ...
''). Cypsela
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
is often used to help determine plant relationships at the genus and species level. The mature
seed A seed is an Plant embryogenesis, embryonic plant enclosed in a testa (botany), protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, includ ...
s usually have little
endosperm The endosperm is a tissue produced inside the seeds of most of the flowering plants following double fertilization. It is triploid (meaning three chromosome sets per nucleus) in most species, which may be auxin-driven. It surrounds the embryo and ...
or none.


Pollen

The pollen of composites is typically echinolophate, a morphological term meaning "with elaborate systems of ridges and spines dispersed around and between the apertures."


Metabolites

In Asteraceae, the energy store is generally in the form of
inulin Inulins are a group of naturally occurring polysaccharides produced by many types of plants, industrially most often extracted from chicory. The inulins belong to a class of dietary fibers known as fructans. Inulin is used by some plants as a me ...
rather than starch. They produce iso/
chlorogenic acid Chlorogenic acid (CGA) is the ester of caffeic acid and (−)-quinic acid, functioning as an intermediate in lignin biosynthesis. The term "chlorogenic acids" refers to a related polyphenol family of esters, including hydroxycinnamic acids (caffe ...
,
sesquiterpene Sesquiterpenes are a class of terpenes that consist of three isoprene units and often have the molecular formula C15H24. Like monoterpenes, sesquiterpenes may be cyclic or contain rings, including many unique combinations. Biochemical modificatio ...
lactones Lactones are cyclic carboxylic esters, containing a 1-oxacycloalkan-2-one structure (), or analogues having unsaturation or heteroatoms replacing one or more carbon atoms of the ring. Lactones are formed by intramolecular esterification of the co ...
, pentacyclic triterpene alcohols, various
alkaloid Alkaloids are a class of basic, naturally occurring organic compounds that contain at least one nitrogen atom. This group also includes some related compounds with neutral and even weakly acidic properties. Some synthetic compounds of similar s ...
s,
acetylene Acetylene (systematic name: ethyne) is the chemical compound with the formula and structure . It is a hydrocarbon and the simplest alkyne. This colorless gas is widely used as a fuel and a chemical building block. It is unstable in its pure f ...
s (cyclic, aromatic, with vinyl end groups),
tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'', ...
s. They have
terpenoid The terpenoids, also known as isoprenoids, are a class of naturally occurring organic chemicals derived from the 5-carbon compound isoprene and its derivatives called terpenes, diterpenes, etc. While sometimes used interchangeably with "terpenes", ...
essential oil An essential oil is a concentrated hydrophobic liquid containing volatile (easily evaporated at normal temperatures) chemical compounds from plants. Essential oils are also known as volatile oils, ethereal oils, aetheroleum, or simply as the o ...
s which never contain
iridoid Iridoids are a type of monoterpenoids in the general form of cyclopentanopyran, found in a wide variety of plants and some animals. They are biosynthetically derived from 8-oxogeranial. Iridoids are typically found in plants as glycosides, mo ...
s. Asteraceae produce
secondary metabolites Secondary metabolites, also called specialised metabolites, toxins, secondary products, or natural products, are organic compounds produced by any lifeform, e.g. bacteria, fungi, animals, or plants, which are not directly involved in the norma ...
, such as
flavonoid Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...
s and
terpenoid The terpenoids, also known as isoprenoids, are a class of naturally occurring organic chemicals derived from the 5-carbon compound isoprene and its derivatives called terpenes, diterpenes, etc. While sometimes used interchangeably with "terpenes", ...
s. Some of these molecules can inhibit protozoan
parasites Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has c ...
such as ''
Plasmodium ''Plasmodium'' is a genus of unicellular eukaryotes that are obligate parasites of vertebrates and insects. The life cycles of ''Plasmodium'' species involve development in a blood-feeding insect host which then injects parasites into a verteb ...
'', ''
Trypanosoma ''Trypanosoma'' is a genus of kinetoplastids (class Trypanosomatidae), a monophyletic group of unicellular parasitic flagellate protozoa. Trypanosoma is part of the phylum Sarcomastigophora. The name is derived from the Greek ''trypano-'' (bore ...
'', ''
Leishmania ''Leishmania'' is a parasitic protozoan, a single-celled organism of the genus ''Leishmania'' that are responsible for the disease leishmaniasis. They are spread by sandflies of the genus ''Phlebotomus'' in the Old World, and of the genus ''Lu ...
'' and parasitic intestinal worms, and thus have potential in medicine.


Taxonomy


History

Compositae, the original name for Asteraceae, were first described in 1740 by Dutch botanist Adriaan van Royen. Traditionally, two subfamilies were recognised:
Asteroideae Asteroideae is a subfamily of the plant family Asteraceae. It contains about 70% of the species of the family. It consists of several tribes, including Astereae, Calenduleae, Eupatorieae, Gnaphalieae, Heliantheae, Senecioneae and Tageteae. As ...
(or Tubuliflorae) and
Cichorioideae The Cichorioideae are a subfamily of the family Asteraceae of flowering plants. Familiar members of Cichorioideae include lettuce, dandelions, chicory and '' Gazania'' species. The subfamily comprises about 240 genera and about 2900 species. It i ...
(or Liguliflorae). The latter has been shown to be extensively
paraphyletic In taxonomy, a group is paraphyletic if it consists of the group's last common ancestor and most of its descendants, excluding a few monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
, and has now been divided into 12 subfamilies, but the former still stands. The study of this family is known as synantherology.


Phylogeny

The
phylogenetic tree A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...
presented below is based on Panero &
Funk Funk is a music genre that originated in African American communities in the mid-1960s when musicians created a rhythmic, danceable new form of music through a mixture of various music genres that were popular among African Americans in the mid ...
(2002) updated in 2014, and now also includes the monotypic
Famatinanthoideae ''Famatinanthus'' is a genus in the family Asteraceae that was described in 2014 and has been assigned to its own tribe Famatinantheae and subfamily Famatinanthoideae. It contains only one known species, ''F. decussatus'', a small shrub of ½—1 ...
. The diamond (♦) denotes a very poorly supported
node In general, a node is a localized swelling (a "knot") or a point of intersection (a vertex). Node may refer to: In mathematics *Vertex (graph theory), a vertex in a mathematical graph *Vertex (geometry), a point where two or more curves, lines, ...
(<50% bootstrap support), the dot (•) a poorly supported node (<80%). The family includes over 32,000 currently accepted species, in over 1,900 genera (
list A ''list'' is any set of items in a row. List or lists may also refer to: People * List (surname) Organizations * List College, an undergraduate division of the Jewish Theological Seminary of America * SC Germania List, German rugby union ...
) in 13 subfamilies. The number of species in the family Asteraceae is rivaled only by Orchidaceae. Which is the larger family is unclear, because of the uncertainty about how many
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extin ...
species each family includes. The four subfamilies
Asteroideae Asteroideae is a subfamily of the plant family Asteraceae. It contains about 70% of the species of the family. It consists of several tribes, including Astereae, Calenduleae, Eupatorieae, Gnaphalieae, Heliantheae, Senecioneae and Tageteae. As ...
,
Cichorioideae The Cichorioideae are a subfamily of the family Asteraceae of flowering plants. Familiar members of Cichorioideae include lettuce, dandelions, chicory and '' Gazania'' species. The subfamily comprises about 240 genera and about 2900 species. It i ...
,
Carduoideae Carduoideae is the thistle subfamily of the Asteraceae, or sunflower family, of flowering plants. It comprises a number of tribes in various circumscriptions of the family, in addition to the Cynareae. Takhtajan, according to Reveal, includes ...
and
Mutisioideae The Mutisioideae are a subfamily in the plant family Asteraceae that includes about 630 species assigned to 44 different genera. This subfamily is mainly native in South America, except for '' Adenocaulon'', '' Chaptalia'', '' Gerbera'', '' Trich ...
contain 99% of the species diversity of the whole family (approximately 70%, 14%, 11% and 3% respectively). Because of the morphological complexity exhibited by this family, agreeing on generic circumscriptions has often been difficult for
taxonomist In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given ...
s. As a result, several of these genera have required multiple revisions.


Paleontology and evolutionary processes

The oldest known fossils of members of Asteraceae are pollen grains from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
of
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
, dated to ∼76–66 mya (
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian ...
to
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
) and assigned to the extant genus ''
Dasyphyllum ''Dasyphyllum'' is a genus of flowering plants in the family Asteraceae. It is distributed in South America, with several species in southeastern Brazil.crown group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. ...
of Asteraceae evolved at least 85.9 mya (Late Cretaceous,
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. T ...
) with a stem node age of 88–89 mya (Late Cretaceous,
Coniacian The Coniacian is an age or stage in the geologic timescale. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series and spans the time between 89.8 ± 1 Ma and 86.3 ± 0.7 Ma (million years ago). The Coniacian is preceded by t ...
). It is not known whether the precise cause of their great success was the development of the highly specialised capitulum, their ability to store energy as
fructan A fructan is a polymer of fructose molecules. Fructans with a short chain length are known as fructooligosaccharides. Fructans can be found in over 12% of the angiosperms including both monocots and dicots such as agave, artichokes, asparagus, leek ...
s (mainly
inulin Inulins are a group of naturally occurring polysaccharides produced by many types of plants, industrially most often extracted from chicory. The inulins belong to a class of dietary fibers known as fructans. Inulin is used by some plants as a me ...
), which is an advantage in relatively dry zones, or some combination of these and possibly other factors. Heterocarpy, or the ability to produce different fruit morphs, has evolved and is common in Asteraceae. It allows seeds to be dispersed over varying distances and each is adapted to different environments, increasing chances of survival.


Etymology and pronunciation

The name Asteraceae () comes to
international scientific vocabulary International scientific vocabulary (ISV) comprises scientific and specialized words whose language of origin may or may not be certain, but which are in current use in several modern languages (that is, translingually, whether in naturalized, lo ...
from
New Latin New Latin (also called Neo-Latin or Modern Latin) is the revival of Literary Latin used in original, scholarly, and scientific works since about 1500. Modern scholarly and technical nomenclature, such as in zoological and botanical taxonomy a ...
, from ''
Aster Aster or ASTER may refer to: Biology * ''Aster'' (genus), a genus of flowering plants ** List of ''Aster'' synonyms, other genera formerly included in ''Aster'' and still called asters in English * Aster (cell biology), a cellular structure shap ...
'', the
type genus In biological taxonomy, the type genus is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearing type of a nominal ...
, + '' -aceae'', a standardized
suffix In linguistics, a suffix is an affix which is placed after the stem of a word. Common examples are case endings, which indicate the grammatical case of nouns, adjectives, and verb endings, which form the conjugation of verbs. Suffixes can carry g ...
for plant family names in modern taxonomy. The genus name comes from the Classical Latin word , "star", which came from Ancient Greek (), "star". It refers to the star-like form of the inflorescence. The original name Compositae is still valid under the International Code of Nomenclature for algae, fungi, and plants. It refers to the "composite" nature of the capitula, which consist of a few or many individual flowers. The vernacular name ''daisy'', widely applied to members of this family, is derived from the
Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlers in the mid-5th c ...
name of the daisy (''
Bellis perennis ''Bellis perennis'' (), the daisy, is a European species of the family Asteraceae, often considered the archetypal species of the name daisy. To distinguish this species from other plants known as daisies, it is sometimes qualified as common dai ...
''): , meaning "day's eye". This is because the petals open at dawn and close at dusk.


Distribution and habitat

Asteraceae species have a widespread distribution, from subpolar to tropical regions in a wide variety of habitats. Most occur in hot desert and cold or hot semi-desert climates, and they are found on every continent but Antarctica. They are especially numerous in tropical and subtropical regions (notably
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. C ...
, eastern Brazil, the
Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the ea ...
, the Levant,
southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number of ...
, central Asia, and
southwestern China Southwest China () is a region in the south of the People's Republic of China. Geography Southwest China is a rugged and mountainous region, transitioning between the Tibetan Plateau to the west and the Chinese coastal hills (东南丘陵) and ...
). The largest proportion of the species occur in the arid and semi-arid regions of subtropical and lower temperate latitudes. The Asteraceae family comprises 10% of all flowering plant species.


Ecology

Asteraceae are especially common in open and dry environments. Many members of Asteraceae are
pollinated Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, a ...
by insects, which explains their value in attracting
beneficial insects Beneficial insects (sometimes called beneficial bugs) are any of a number of species of insects that perform valued services like pollination and pest control. The concept of ''beneficial'' is subjective and only arises in light of desired outcome ...
, but
anemophily Anemophily or wind pollination is a form of pollination whereby pollen is distributed by wind. Almost all gymnosperms are anemophilous, as are many plants in the order Poales, including grasses, sedges, and rushes. Other common anemophilous plan ...
is also present (e.g. ''
Ambrosia In the ancient Greek myths, ''ambrosia'' (, grc, ἀμβροσία 'immortality'), the food or drink of the Greek gods, is often depicted as conferring longevity or immortality upon whoever consumed it. It was brought to the gods in Olympus by ...
'', '' Artemisia''). There are many
apomictic In botany, apomixis is asexual reproduction without fertilization. Its etymology is Greek for "away from" + "mixing". This definition notably does not mention meiosis. Thus "normal asexual reproduction" of plants, such as propagation from cutt ...
species in the family. Seeds are ordinarily dispersed intact with the fruiting body, the cypsela. ''
Anemochory In Spermatophyte plants, seed dispersal is the movement, spread or transport of seeds away from the parent plant. Plants have limited mobility and rely upon a variety of dispersal vectors to transport their seeds, including both abiotic vectors, ...
'' (wind dispersal) is common, assisted by a hairy pappus. ''
Epizoochory In Spermatophyte plants, seed dispersal is the movement, spread or transport of seeds away from the parent plant. Plants have limited mobility and rely upon a variety of dispersal vectors to transport their seeds, including both abiotic vectors, ...
'' is another common method, in which the dispersal unit, a single cypsela (e.g. ''
Bidens ''Bidens'' is a genus of flowering plants in the aster family, Asteraceae.''Bidens''.
Flo ...
'') or entire capitulum (e.g. ''
Arctium ''Arctium'' is a genus of biennial plants commonly known as burdock, family Asteraceae. Native to Europe and Asia, several species have been widely introduced worldwide. Burdock's clinging properties, in addition to providing an excellent mecha ...
'') has hooks, spines or some structure to attach to the fur or plumage (or even clothes, as in the photo) of an animal just to fall off later far from its mother plant. Some members of Asteraceae are economically important as weeds. Notable in the United States are '' Senecio jacobaea'' (ragwort), ''
Senecio vulgaris ''Senecio vulgaris'', often known by the common names groundsel and old-man-in-the-spring, is a flowering plant in the family Asteraceae. It is an annual herb, native to Europe and widely naturalised as a ruderal species in suitable disturbed ha ...
'' (groundsel), and ''
Taraxacum ''Taraxacum'' () is a large genus of flowering plants in the family Asteraceae, which consists of species commonly known as dandelions. The scientific and hobby study of the genus is known as taraxacology. The genus is native to Eurasia and Nor ...
'' (dandelion). Some are
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
in particular regions, often having been introduced by human agency. Examples include various
tumbleweed A tumbleweed is a structural part of the above-ground anatomy of a number of species of plants. It is a diaspore that, once mature and dry, detaches from its root or stem and rolls due to the force of the wind. In most such species, the tumbl ...
s, ''
Bidens ''Bidens'' is a genus of flowering plants in the aster family, Asteraceae.''Bidens''.
Flo ...
'', ragweeds,
thistle Thistle is the common name of a group of flowering plants characterised by leaves with sharp prickles on the margins, mostly in the family Asteraceae. Prickles can also occur all over the planton the stem and on the flat parts of the leaves. ...
s, and
dandelion ''Taraxacum'' () is a large genus of flowering plants in the family Asteraceae, which consists of species commonly known as dandelions. The scientific and hobby study of the genus is known as taraxacology. The genus is native to Eurasia and Nor ...
. Dandelion was introduced into
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the C ...
by European settlers who used the young leaves as a salad green. A number of species are toxic to grazing animals.


Uses

Asteraceae is an economically important family, providing products such as
cooking oil Cooking oil is plant, animal, or synthetic liquid fat used in frying, baking, and other types of cooking. It is also used in food preparation and flavoring not involving heat, such as salad dressings and bread dips, and may be called edible oil ...
s,
leaf vegetable Leaf vegetables, also called leafy greens, pot herbs, vegetable greens, or simply greens, are plant leaves eaten as a vegetable, sometimes accompanied by tender petioles and shoots. Leaf vegetables eaten raw in a salad can be called salad gre ...
s like
lettuce Lettuce (''Lactuca sativa'') is an annual plant of the family Asteraceae. It is most often grown as a leaf vegetable, but sometimes for its stem and seeds. Lettuce is most often used for salads, although it is also seen in other kinds of food, ...
,
sunflower seed The sunflower seed is the seed of the sunflower (''Helianthus annuus''). There are three types of commonly used sunflower seeds: linoleic (most common), high oleic, and sunflower oil seeds. Each variety has its own unique levels of monounsat ...
s,
artichoke The globe artichoke (''Cynara cardunculus'' var. ''scolymus'' ),Rottenberg, A., and D. Zohary, 1996: "The wild ancestry of the cultivated artichoke." Genet. Res. Crop Evol. 43, 53–58. also known by the names French artichoke and green articho ...
s, sweetening agents, coffee substitutes and
herbal tea Herbal teas, also known as herbal infusions and less commonly called tisanes (UK and US , US also ), are beverages made from the infusion or decoction of herbs, spices, or other plant material in hot water. Oftentimes herb tea, or the plain term ...
s. Several genera are of
horticultural Horticulture is the branch of agriculture that deals with the art, science, technology, and business of plant cultivation. It includes the cultivation of fruits, vegetables, nuts, seeds, herbs, sprouts, mushrooms, algae, flowers, seaweeds and no ...
importance, including pot marigold (''
Calendula officinalis ''Calendula officinalis'', the pot marigold, common marigold, ruddles, Mary's gold or Scotch marigold, is a flowering plant in the daisy family Asteraceae. It is probably native to southern Europe, though its long history of cultivation makes its ...
''), ''
Echinacea ''Echinacea'' is a genus of herbaceous flowering plants in the daisy family. It has ten species, which are commonly called coneflowers. They are found only in eastern and central North America, where they grow in moist to dry prairies and ope ...
'' (coneflowers), various daisies,
fleabane Fleabane is a common name for some flowering plants in the family Asteraceae. Most are in the subfamily Asteroideae: * ''Conyza'' (butterweeds or horseweeds: Astereae) * ''Erigeron'' (Astereae) * '' Inula'' ("yellowheads": Inuleae) * '' Pluchea' ...
,
chrysanthemum Chrysanthemums (), sometimes called mums or chrysanths, are flowering plants of the genus ''Chrysanthemum'' in the family Asteraceae. They are native to East Asia and northeastern Europe. Most species originate from East Asia and the center o ...
s,
dahlia Dahlia (, ) is a genus of bushy, tuberous, herbaceous perennial plants native to Mexico and Central America. A member of the Asteraceae (former name: Compositae) family of dicotyledonous plants, its garden relatives thus include the sunflower, ...
s,
zinnia ''Zinnia'' is a genus of plants of the tribe Heliantheae within the family Asteraceae. They are native to scrub and dry grassland in an area stretching from the Southwestern United States to South America, with a centre of diversity in Mexico. ...
s, and
helenium ''Helenium'' is a genus of annuals and herbaceous perennial plants in the family Asteraceae, native to the Americas. They bear yellow or orange daisy-like composite flowers. A number of these species (particularly '' Helenium autumnale'') have ...
s. Asteraceae are important in herbal medicine, including '' Grindelia'',
yarrow ''Achillea millefolium'', commonly known as yarrow () or common yarrow, is a flowering plant in the family Asteraceae. Other common names include old man's pepper, devil's nettle, sanguinary, milfoil, soldier's woundwort, and thousand seal. T ...
, and many others. Commercially important plants in Asteraceae include the food crops ''
Lactuca sativa Lettuce (''Lactuca sativa'') is an annual plant of the family Asteraceae. It is most often grown as a leaf vegetable, but sometimes for its stem and seeds. Lettuce is most often used for salads, although it is also seen in other kinds of food, ...
'' (lettuce), '' Cichorium'' (chicory), ''
Cynara scolymus The globe artichoke (''Cynara cardunculus'' var. ''scolymus'' ),Rottenberg, A., and D. Zohary, 1996: "The wild ancestry of the cultivated artichoke." Genet. Res. Crop Evol. 43, 53–58. also known by the names French artichoke and green articho ...
'' (globe artichoke), ''
Helianthus annuus The common sunflower (''Helianthus annuus'') is a large annual forb of the genus ''Helianthus'' grown as a crop for its edible oily seeds. Apart from cooking oil production, it is also used as livestock forage (as a meal or a silage plant), as b ...
'' (sunflower), '' Smallanthus sonchifolius'' (yacón), ''
Carthamus tinctorius Safflower (''Carthamus tinctorius'') is a highly branched, herbaceous, thistle-like annual plant in the family Asteraceae. It is commercially cultivated for vegetable oil extracted from the seeds and was used by the early Spanish colonies along ...
'' (safflower) and '' Helianthus tuberosus'' (Jerusalem artichoke). Plants are used as herbs and in
herbal tea Herbal teas, also known as herbal infusions and less commonly called tisanes (UK and US , US also ), are beverages made from the infusion or decoction of herbs, spices, or other plant material in hot water. Oftentimes herb tea, or the plain term ...
s and other beverages. Chamomile, for example, comes from two different species: the annual ''
Matricaria chamomilla ''Matricaria chamomilla'' (synonym: ''Matricaria recutita''), commonly known as chamomile (also spelled camomile), German chamomile, Hungarian chamomile (kamilla), wild chamomile, blue chamomile, or scented mayweed, is an annual plant of the com ...
'' (German chamomile) and the perennial ''
Chamaemelum nobile ''Chamaemelum nobile'', commonly known as chamomile (also spelled camomile), is a low perennial plant found in dry fields and around gardens and cultivated grounds in Europe, North America, and South America. Its synonym is ''Anthemis nobili ...
'' (Roman chamomile). ''
Calendula ''Calendula'' () is a genus of about 15–20 species''Calendula''.
Flora of China.
'' (known as pot marigold) is grown commercially for herbal teas and
potpourri Potpourri ( ) is a mixture of dried, naturally fragrant plant materials used to provide a gentle natural scent, commonly in residential settings. It is often placed in a decorative bowl. The word "potpourri" comes into English from the French ...
. ''
Echinacea ''Echinacea'' is a genus of herbaceous flowering plants in the daisy family. It has ten species, which are commonly called coneflowers. They are found only in eastern and central North America, where they grow in moist to dry prairies and ope ...
'' is used as a medicinal tea. The wormwood genus '' Artemisia'' includes
absinthe Absinthe (, ) is an anise-flavoured spirit derived from several plants, including the flowers and leaves of ''Artemisia absinthium'' ("grand wormwood"), together with green anise, sweet fennel, and other medicinal and culinary herbs. Historical ...
(''A. absinthium'') and
tarragon Tarragon (''Artemisia dracunculus''), also known as estragon, is a species of perennial herb in the family Asteraceae. It is widespread in the wild across much of Eurasia and North America and is cultivated for culinary and medicinal purposes. O ...
(''A. dracunculus''). Winter tarragon (''
Tagetes lucida ''Tagetes lucida'' is a perennial plant native to Mexico and Central America. It is used as a medicinal plant and as a culinary herb. The leaves have a tarragon-like scent, with hints of anise, and it has entered the nursery trade in North A ...
''), is commonly grown and used as a tarragon substitute in climates where tarragon will not survive. Many members of the family are grown as ornamental plants for their flowers, and some are important ornamental crops for the cut flower industry. Some examples are ''
Chrysanthemum Chrysanthemums (), sometimes called mums or chrysanths, are flowering plants of the genus ''Chrysanthemum'' in the family Asteraceae. They are native to East Asia and northeastern Europe. Most species originate from East Asia and the center o ...
'', ''
Gerbera ''Gerbera'' ( or ) L. is a genus of plants in the Asteraceae (Compositae) family. The first scientific description of a ''Gerbera'' was made by J. D. Hooker in Curtis's Botanical Magazine in 1889 when he described '' Gerbera jamesonii'', a Sou ...
'', ''
Calendula ''Calendula'' () is a genus of about 15–20 species''Calendula''.
Flora of China.
'', '' Dendranthema'', '' Argyranthemum'', ''
Dahlia Dahlia (, ) is a genus of bushy, tuberous, herbaceous perennial plants native to Mexico and Central America. A member of the Asteraceae (former name: Compositae) family of dicotyledonous plants, its garden relatives thus include the sunflower, ...
'', ''
Tagetes ''Tagetes'' () is a genusSoule, J. A. 1996. Infrageneric Systematics of Tagetes. Pgs. 435-443 in Compositae: Systematics, Proceedings of the International Compositae Conference, Kew 1994, Vol. I, Eds. D.J.N. Hind & H.J. Beentje. of annual or ...
'', ''
Zinnia ''Zinnia'' is a genus of plants of the tribe Heliantheae within the family Asteraceae. They are native to scrub and dry grassland in an area stretching from the Southwestern United States to South America, with a centre of diversity in Mexico. ...
'', and many others. Many species of this family possess medicinal properties and are used as traditional
antiparasitic Antiparasitics are a class of medications which are indicated for the treatment of parasitic diseases, such as those caused by helminths, amoeba, ectoparasites, parasitic fungi, and protozoa, among others. Antiparasitics target the parasitic agent ...
medicine. Members of the family are also commonly featured in medical and phytochemical journals because the
sesquiterpene Sesquiterpenes are a class of terpenes that consist of three isoprene units and often have the molecular formula C15H24. Like monoterpenes, sesquiterpenes may be cyclic or contain rings, including many unique combinations. Biochemical modificatio ...
lactone Lactones are cyclic carboxylic esters, containing a 1-oxacycloalkan-2-one structure (), or analogues having unsaturation or heteroatoms replacing one or more carbon atoms of the ring. Lactones are formed by intramolecular esterification of the co ...
compounds contained within them are an important cause of
allergic contact dermatitis Allergic contact dermatitis (ACD) is a form of contact dermatitis that is the manifestation of an allergic response caused by contact with a substance; the other type being irritant contact dermatitis (ICD). Although less common than ICD, ACD is ...
. Allergy to these compounds is the leading cause of allergic contact dermatitis in florists in the US.
Pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophy ...
from ragweed ''
Ambrosia In the ancient Greek myths, ''ambrosia'' (, grc, ἀμβροσία 'immortality'), the food or drink of the Greek gods, is often depicted as conferring longevity or immortality upon whoever consumed it. It was brought to the gods in Olympus by ...
'' is among the main causes of so-called
hay fever Allergic rhinitis, of which the seasonal type is called hay fever, is a type of inflammation in the nose that occurs when the immune system overreacts to allergens in the air. Signs and symptoms include a runny or stuffy nose, sneezing, red, i ...
in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territo ...
. Asteraceae are also used for some industrial purposes. French Marigold (''
Tagetes patula ''Tagetes patula'', the French marigold, is a species of flowering plant in the family Asteraceae, native to Mexico and Guatemala with several naturalised populations in many other countries. It is widely cultivated as an easily grown bedding pl ...
'') is common in commercial poultry feeds and its oil is extracted for uses in cola and the cigarette industry. The genera ''
Chrysanthemum Chrysanthemums (), sometimes called mums or chrysanths, are flowering plants of the genus ''Chrysanthemum'' in the family Asteraceae. They are native to East Asia and northeastern Europe. Most species originate from East Asia and the center o ...
'', ''
Pulicaria ''Pulicaria'' is a genus of flowering plant in the sunflower family, native to Europe, Asia, and Africa. In North America ''Pulicaria'' is known by the common name false fleabane. ''Pulicaria'' species accepted by the Plants of the World Onli ...
'', ''
Tagetes ''Tagetes'' () is a genusSoule, J. A. 1996. Infrageneric Systematics of Tagetes. Pgs. 435-443 in Compositae: Systematics, Proceedings of the International Compositae Conference, Kew 1994, Vol. I, Eds. D.J.N. Hind & H.J. Beentje. of annual or ...
'', and ''
Tanacetum ''Tanacetum'' is a genus of about 160 species of flowering plants in the aster family, Asteraceae, native to many regions of the Northern Hemisphere.
'' contain species with useful insecticidal properties. ''
Parthenium argentatum ''Parthenium argentatum'', commonly known as the guayule ( or , as in Spanish), is a perennial woody shrub in the family Asteraceae that is native to the rangeland area of the Chihuahuan Desert; including the southwestern United States and north ...
'' (guayule) is a source of
hypoallergenic Hypoallergenic, meaning "below average" or "slightly" allergenic, is a term meaning that something (usually cosmetics, pets, textiles, food, etc.) causes fewer allergic reactions. The term was first used in 1953 in an advertising campaign for cos ...
latex Latex is an emulsion (stable dispersion) of polymer microparticles in water. Latexes are found in nature, but synthetic latexes are common as well. In nature, latex is found as a milky fluid found in 10% of all flowering plants (angiosperms ...
. Several members of the family are copious
nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualis ...
producers and are useful for evaluating
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the maj ...
populations during their bloom. ''
Centaurea ''Centaurea'' () is a genus of over 700 species of herbaceous thistle-like flowering plants in the family Asteraceae. Members of the genus are found only north of the equator, mostly in the Eastern Hemisphere; the Middle East and surrounding reg ...
'' (knapweed), ''Helianthus annuus'' (domestic sunflower), and some species of ''
Solidago ''Solidago'', commonly called goldenrods, is a genus of about 100 to 120''Solidago''.
F ...
'' (goldenrod) are major "
honey plant Honeybees usually collect nectar, pollen, or both from the following species of plants, which are called honey plants, for making honey. Acanthaceae (Acanthus family) *'' Avicennia nitida'' Jacq. or Avicennia ge ...
s" for
beekeeper A beekeeper is a person who keeps honey bees. Beekeepers are also called honey farmers, apiarists, or less commonly, apiculturists (both from the Latin '' apis'', bee; cf. apiary). The term beekeeper refers to a person who keeps honey bees i ...
s. ''Solidago'' produces relatively high protein
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophy ...
, which helps
honey bee A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the current cosmop ...
s over winter.


References


Bibliography

*


External links

* *
Asteraceae
at th
''Angiosperm Phylogeny Website''

Compositae.org
– Compositae Working Group (CWG) and Global Compositae Database (GCD) {{Authority control Asterales families Extant Campanian first appearances