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The coconut tree (''Cocos nucifera'') is a member of the palm tree
family In human society, family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of families is to maintain the well-being of its members an ...
( Arecaceae) and the only living
species In biology, a species is the basic unit of biological classification, classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individu ...
of the
genus Genus (plural genera) is a taxonomic rank Taxonomy (general) is the practice and science of classification of things or concepts, including the principles that underlie such classification. The term may also refer to a specific classification s ...
''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, the
seed A seed is an embryonic plant enclosed in a protective outer covering. The formation of the seed is part of the process of reproduction Reproduction (or procreation or breeding) is the biological process by which new individual organism ...
, or the
fruit In botany, a fruit is the seed-bearing structure in flowering plants (also known as angiosperms) formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate seeds. Edible fruits, in particular, have propa ...
, which botanically is a drupe, not a nut. The name comes from the old Portuguese word '' coco'', meaning "head" or "skull", after the three indentations on the coconut shell that resemble facial features. They are ubiquitous in coastal tropical regions and are a cultural icon of the
tropics The tropics are the region of Earth surrounding the Equator. They are delimited in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S; these latitudes correspond to the ...
. It is one of the most useful trees in the world and is often referred to as the "tree of life". It provides food, fuel, cosmetics, folk medicine and building materials, among many other uses. The inner flesh of the mature seed, as well as the coconut milk extracted from it, form a regular part of the diets of many people in the
tropics The tropics are the region of Earth surrounding the Equator. They are delimited in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S; these latitudes correspond to the ...
and
subtropics The subtropical zones or subtropics are geographical zone, geographic and Köppen climate classification, climate zones located to the north and south of the tropics, tropical zone. Geographically part of the north and south temperate zones, th ...
. Coconuts are distinct from other fruits because their endosperm contains a large quantity of clear liquid, called '' coconut water'' or ''coconut juice''. Mature, ripe coconuts can be used as edible seeds, or processed for
oil An oil is any nonpolar chemical substance A chemical substance is a form of matter In classical physics and general chemistry, matter is any substance that has mass and takes up space by having volume. All everyday objects that can b ...
and plant milk from the flesh,
charcoal Charcoal is a lightweight black carbon Carbon (from la, carbo "coal") is a chemical element Image:Simple Periodic Table Chart-blocks.svg, 400px, Periodic table, The periodic table of the chemical elements In chemistry, an elemen ...

charcoal
from the hard shell, and coir from the fibrous husk. Dried coconut flesh is called copra, and the oil and milk derived from it are commonly used in cooking – frying in particular – as well as in soaps and cosmetics. The hard shells, fibrous husks and long pinnate leaves can be used as material to make a variety of products for furnishing and decoration. The coconut has cultural and religious significance in certain societies, particularly in India, where it is used in Hindu rituals. It forms the basis of wedding and worship rituals in Hinduism, a Coconut Religion, coconut religion in Vietnam, and features in the Origin myth, origin myths of several societies. The falling nature of their mature fruit has led to preoccupation with death by coconut. Coconuts have been used by humans for thousands of years and may have spread to their present range by Pacific island settlers. The evolutionary origin of the coconut is under dispute, with theories stating that it may have evolved in Asia, South America, or on Pacific islands. Trees grow up to tall and can yield up to 75 fruits per year, though fewer than 30 is more typical. Plants are intolerant of cold weather and prefer copious precipitation, as well as full sunlight. Many insect pests and diseases affect the species and are a nuisance for commercial production. About 75% of the world's supply of coconuts is produced by Indonesia, the Philippines, and India combined.


Etymology

The name ''coconut'' is derived from the 16th-century Portuguese language, Portuguese word '' coco'', meaning 'head' or 'skull' after the three indentations on the coconut shell that resemble facial features. ''Coco'' and ''coconut'' apparently came from 1521 encounters by Portuguese and Spanish people, Spanish explorers with Pacific islanders, with the coconut shell reminding them of a ghost or witch in Portuguese folklore called '' coco'' (also ''côca'').Losada, Fernando Díez. (2004). ''La tribuna del idioma''. Editorial Tecnologica de CR
p. 481
. .
In the West it was originally called ''nux indica'', a name used by Marco Polo in 1280 while in Sumatra. He took the term from the Arabs, who called it جوز هندي ''jawz hindī'', translating to 'Indian nut'.Elzebroek, A.T.G. and Koop Wind (Eds.). (2008). ''Guide to Cultivated Plants''. CABI. pp
186–192
. .
''Thenga'', its Malayalam name, was used in the detailed description of coconut found in ''Itinerario'' by Ludovico di Varthema published in 1510 and also in the later ''Hortus Indicus Malabaricus''.Grimwood 1975, p
1
.
The specific name (botany), specific name ''nucifera'' is derived from the Latin words ''nux'' (nut) and ''fera'' (bearing), for 'nut-bearing'.


History


Origin

The American botanist Orator F. Cook proposed a theory in 1901 on the location of the origin of ''Cocos nucifera'' based on its current worldwide distribution. He hypothesized that the coconut originated in the Americas, based on his belief that American coconut populations predated European contact and because he considered pan-tropical distribution by ocean currents improbable. Modern genetic studies have identified the center of origin of coconuts as being the region between Southwest Asia and Melanesia, where it shows greatest genetic diversity.Perera, Lalith, Suriya A.C.N. Perera, Champa K. Bandaranayake and Hugh C. Harries. (2009). "Chapter 12 – Coconut". In Johann Vollmann and Istvan Rajcan (Eds.). ''Oil Crops''. Springer. pp
370–372
. .
Their cultivation and spread was closely tied to the early migrations of the Austronesian peoples who carried coconuts as canoe plants to islands they settled. The similarities of the local names in the Austronesian region is also cited as evidence that the plant originated in the region. For example, the Polynesian languages, Polynesian and Melanesian languages, Melanesian term ''niu''; Tagalog language, Tagalog and Chamorro language, Chamorro term ''niyog''; and the Malay language, Malay word ''nyiur'' or ''nyior''. A study in 2011 identified two highly genetically differentiated subpopulations of coconuts, one originating from Island Southeast Asia (the Pacific group) and the other from the southern margins of the Indian subcontinent (the Indo-Atlantic group). The Pacific group is the only one to display clear genetic and phenotypic indications that they were domesticated; including dwarf habit, self-pollination, and the round "''niu vai''" fruit morphology with larger endosperm-to-husk ratios. The distribution of the Pacific coconuts correspond to the regions settled by Austronesian voyagers indicating that its spread was largely the result of human introductions. It is most strikingly displayed in Madagascar, an island settled by Austronesian sailors at around 2000 to 1500 Before present, BP. The coconut populations in the island show genetic admixture between the two subpopulations indicating that Pacific coconuts were brought by the Austronesian settlers that later interbred with the local Indo-Atlantic coconuts. Genetic studies of coconuts have also confirmed pre-Columbian populations of coconuts in Panama in South America. However, it is not native and display a genetic bottleneck resulting from a founder effect. A study in 2008 showed that the coconuts in the Americas are genetically closest related to coconuts in the Philippines, and not to any other nearby coconut populations (including Polynesia). Such an origin indicates that the coconuts were not introduced naturally, such as by sea currents. The researchers concluded that it was brought by early Austronesian sailors to the Americas from at least 2,250 BP, and may be proof of pre-Columbian contact between Austronesian cultures and South American cultures, albeit in the opposite direction than what early hypotheses like Heyerdahl's had proposed. It is further strengthened by other similar botanical evidence of contact, like the pre-colonial presence of sweet potato in Oceanian cultures. During the Early modern period, colonial era, Pacific coconuts were further introduced to Mexico from the Spanish East Indies via the Manila galleons. In contrast to the Pacific coconuts, Indo-Atlantic coconuts were largely spread by Arab and Persian traders into the East African coast. Indo-Atlantic coconuts were also introduced into the Atlantic Ocean by Portuguese ships from their colonies in coastal India and Sri Lanka; first being introduced to coastal West Africa, then onwards into the Caribbean and the east coast of Brazil. All of these introductions are within the last few centuries, relatively recent in comparison to the spread of Pacific coconuts. In attempting to determine whether the species had originated in South America or Asia, a 2014 study proposed that it was neither, and that the species evolved on coral atolls in the Pacific. Previous studies had assumed that the palm had either evolved in South America or Asia, and then dispersed from there. The 2014 study hypothesized that instead the species evolved while on coral atolls in the Pacific, and then dispersed to the continents. It contended that this would have provided the necessary evolutionary pressures, and would account for morphological factors such as a thick husk to protect against ocean degradation and provide a moist medium in which to germinate on sparse atolls.


Evolutionary history

The evolutionary history and fossil distribution of ''Cocos nucifera'' and other members of the tribe Cocoseae is more ambiguous than modern-day dispersal and distribution, with its ultimate origin and pre-human dispersal still unclear. There are currently two major viewpoints on the origins of the genus ''Cocos'', one in the Indo-Pacific, and another in South America. The vast majority of ''Cocos''-like fossils have been recovered generally from only two regions in the world: New Zealand and west-central India. However, like most palm fossils, ''Cocos''-like fossils are still putative, as they are usually difficult to identify. The earliest ''Cocos''-like fossil to be found was ''"Cocos" zeylanica'', a fossil species described from small fruits, around × in size, recovered from the Miocene (~23 to 5.3 million years ago) of New Zealand in 1926. Since then, numerous other fossils of similar fruits were recovered throughout New Zealand from the Eocene, Oligocene, and possibly the Holocene. But research on them is still ongoing to determine which of them (if any) actually belong to the genus ''Cocos''. Endt & Hayward (1997) have noted their resemblance to members of the South American genus ''Parajubaea'', rather than ''Cocos'', and propose a South American origin. Conran ''et al.'' (2015), however, suggests that their diversity in New Zealand indicate that they evolved endemically, rather than being introduced to the islands by long-distance dispersal. In west-central India, numerous fossils of ''Cocos''-like fruits, leaves, and stems have been recovered from the Deccan Traps. They include morphotaxa like ''Palmoxylon sundaran'', ''Palmoxylon insignae'', and ''Palmocarpon cocoides''. ''Cocos''-like fossils of fruits include ''"Cocos" intertrappeansis'', ''"Cocos" pantii'', and ''"Cocos" sahnii''. They also include fossil fruits that have been tentatively identified as modern ''Cocos nucifera''. These includes two specimens named ''"Cocos" palaeonucifera'' and ''"Cocos" binoriensis'', both were dated by their authors to the Maastrichtian–Danian of the early Tertiary (70 to 62 million years ago). ''C. binoriensis'' has been claimed by their authors to be the earliest known fossil of ''Cocos nucifera''. Outside of New Zealand and India, only two other regions have reported ''Cocos''-like fossils, namely Australia and Colombia. In Australia, a ''Cocos''-like fossil fruit, measuring , were recovered from the Chinchilla Sand Formation dated to the latest Pliocene or basal Pleistocene. Rigby (1995) assigned them to modern ''Cocos nucifera'' based on its size. In Colombia, a single ''Cocos''-like fruit was recovered from the Paleocene, middle to late Paleocene Cerrejón Formation. The fruit however was compacted in the fossilization process and it was not possible to determine if it had the diagnostic three pores that characterize members of the tribe Cocoseae. Nevertheless, the authors Gomez-Navarro ''et al.'' (2009), assigned it to ''Cocos'' based on the size and the ridged shape of the fruit.


Historical records

Literary evidence from the ''Ramayana'' and History of Sri Lanka, Sri Lankan chronicles indicates that the coconut was present in the Indian subcontinent before the 1st century BCE. The earliest direct description is given by Cosmas Indicopleustes in his ''Topographia Christiana'' written around 545, referred to as "the great nut of India".Rosengarten, Frederic Jr. (2004). ''The Book of Edible Nuts''. Dover Publications. pp
65–93
. .
Another early mention of the coconut dates back to the "One Thousand and One Nights" story of Sinbad the Sailor wherein he bought and sold a coconut during his fifth voyage. In March 1521, a description of the coconut was given by Antonio Pigafetta writing in Italian and using the words "''cocho''"/"''cochi''", as recorded in his journal after the first European crossing of the Pacific Ocean during the Ferdinand Magellan, Magellan circumnavigation and meeting the inhabitants of what would become known as Guam and the Philippines. He explained how at Guam "they eat coconuts" ("''mangiano cochi''") and that the natives there also "anoint the body and the hair with coconut and sesame seed, beniseed oil" ("''ongieno el corpo et li capili co oleo de cocho et de giongioli''").


Description


Plant

''Cocos nucifera'' is a large palm, growing up to tall, with pinnate leaves long, and pinnae long; old leaves break away cleanly, leaving the trunk (botany), trunk smooth.T. Pradeepkumar, B. Sumajyothibhaskar, and K.N. Satheesan. (2008). ''Management of Horticultural Crops'' (Horticulture Science Series Vol.11, 2nd of 2 Parts). New India Publishing. pp
539–587
. .
On fertile soil, a tall coconut palm tree can yield up to 75 fruits per year, but more often yields less than 30.Sarian, Zac B. (August 18, 2010)
New coconut yields high
. ''The Manila Bulletin''. Retrieved April 21, 2011.
Ravi, Rajesh. (March 16, 2009)
Rise in coconut yield, farming area put India on top
. ''The Financial Express''. Retrieved April 21, 2011.
Given proper care and growing conditions, coconut palms produce their first fruit in six to ten years, taking 15 to 20 years to reach peak production. Many different varieties are grown, including the Maypan coconut palm, Maypan coconut, King coconut, and Macapuno. These vary by the taste of the coconut water and color of the fruit, as well as other genetic factors. Dwarf plant, Dwarf varieties are also available.


Fruit

Botany, Botanically, the coconut
fruit In botany, a fruit is the seed-bearing structure in flowering plants (also known as angiosperms) formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate seeds. Edible fruits, in particular, have propa ...
is a drupe, not a true nut. Like other fruits, it has Fruit anatomy, three layers: the exocarp, mesocarp, and endocarp. The exocarp and mesocarp make up the "husk" of the coconuts. The endosperm is initially in its nuclear phase suspended within the coconut water. As development continues, cellular layers of endosperm deposit along the walls of the coconut, becoming the edible coconut "flesh". Coconuts sold in the shops of nontropical countries often have had the exocarp (outermost layer) removed. The mesocarp is composed of a fiber, called coir, which has many traditional and commercial uses. The shell has three germination pores (Micropyle (botany), micropyles) or "eyes" that are clearly visible on its outside surface once the husk is removed. A full-sized coconut weighs about . It takes around 6,000 full-grown coconuts to produce one tonne of copra.


Roots

Unlike some other plants, the palm tree has neither a tap root nor root hairs, but has a fibrous root system.Thampan, P.K. (1981). ''Handbook on Coconut Palm''. Oxford & IBH Publishing Co. The root system consists of an abundance of thin roots that grow outward from the plant near the surface. Only a few of the roots penetrate deep into the soil for stability. This type of root system is known as fibrous or adventitious, and is a characteristic of grass species. Other types of large trees produce a single downward-growing tap root with a number of feeder roots growing from it. 2,000-4,000 Adventitious Root, adventitious roots may grow, each about large. Decayed roots are replaced regularly as the tree grows new ones.


Inflorescence

The palm produces both the female and male flowers on the same inflorescence; thus, the palm is monoecious. However, there is some evidence that it may be polygamomonoecious, and may occasionally have bisexual flowers. The female flower is much larger than the male flower. Flowering occurs continuously. Coconut palms are believed to be largely cross-pollination, pollinated, although most dwarf varieties are self-pollinating.


Distribution

Coconuts have a nearly cosmopolitan distribution thanks to human action in using them for agriculture. However their historical distribution was likely more limited.


Natural habitat

The coconut palm thrives on sandy soils and is highly tolerant of salinity. It prefers areas with abundant sunlight and regular rainfall ( annually), which makes colonizing shorelines of the tropics relatively straightforward.Chan, Edward and Craig R. Elevitch. (April 2006)
''Cocos nucifera'' (coconut)
(version 2.1). In C. R. Elevitch (ed.). ''Species Profiles for Pacific Island Agroforestry''. Hōlualoa, Hawai‘i: Permanent Agriculture Resources (PAR).
Coconuts also need high humidity (at least 70–80%) for optimum growth, which is why they are rarely seen in areas with low humidity. However, they can be found in humid areas with low annual precipitation such as in Karachi, Pakistan, which receives only about of rainfall per year, but is consistently warm and humid. Coconut palms require warm conditions for successful growth, and are intolerant of cold weather. Some seasonal variation is tolerated, with good growth where mean summer temperatures are between , and survival as long as winter temperatures are above ; they will survive brief drops to . Severe frost is usually fatal, although they have been known to recover from temperatures of . They may grow but not fruit properly in areas with insufficient warmth, such as Bermuda. The conditions required for coconut trees to grow without any care are: * Mean daily temperature above every day of the year * Mean annual rainfall above * No or very little overhead Canopy (biology), canopy, since even small trees require direct sun The main limiting factor for most locations which satisfy the rainfall and temperature requirements is canopy growth, except those locations near coastlines, where the sandy soil and salt spray limit the growth of most other trees.


Domestication

Coconuts could not reach inland locations without human intervention (to carry seednuts, plant seedlings, etc.) and early germination on the palm (vivipary) was important, rather than increasing the number or size of the edible parts of a fruit that was already large enough. Human cultivation of the coconut selected, not for larger size, but for thinner husks and increased volume of endosperm, the solid "meat" or liquid "water" that provides the fruit its food value. Although these modifications for domestication would reduce the fruit's ability to float, this ability would be irrelevant to a cultivated population. Among modern ''C. nucifera'', two major types or variants occur: a thick-husked, angular fruit and a thin-husked, spherical fruit with a higher proportion of endosperm reflect a trend of cultivation in ''C. nucifera''. The first coconuts were of the ''niu kafa'' type, with thick husks to protect the seed, an angular, highly ridged shape to promote buoyancy during ocean dispersal, and a pointed base that allowed fruits to dig into the sand, preventing them from being washed away during germination on a new island. As early human communities began to harvest coconuts for eating and planting, they (perhaps unintentionally) selected for a larger endosperm-to-husk ratio and a broader, spherical base, which rendered the fruit useful as a cup or bowl, thus creating the ''niu vai'' type. The decreased buoyancy and increased fragility of this spherical, thin-husked fruit would not matter for a species that had started to be dispersed by humans and grown in plantations. Harries' adoption of the Polynesian terms ''niu kafa'' and ''niu vai'' has now passed into general scientific discourse, and his hypothesis is generally accepted. Variants of ''C. nucifera'' are also categorized as tall (var. ''typica'') or dwarf (var. ''nana''). The two groups are genetically distinct, with the dwarf variety showing a greater degree of artificial selection for ornamental traits and for early germination and fruiting. The tall variety is outcrossing while dwarf palms are incrossing, which has led to a much greater degree of genetic diversity within the tall group. The dwarf subspecies is thought to have mutated from the tall group under human selection pressure.


Dispersal

Coconut fruit in the wild are light, buoyant, and highly water resistant. It is claimed that they evolved to disperse significant distances via marine currents.Foale, Mike. (2003)
The Coconut Odyssey – the bounteous possibilities of the tree of life

Australian Centre for International Agricultural Research
. Retrieved May 30, 2009.
However, it can also be argued that the placement of the vulnerable eye of the nut (down when floating), and the site of the coir cushion are better positioned to ensure that the water-filled nut does not fracture when dropping on rocky ground, rather than for flotation. It is also often stated that coconuts can travel 110 days, or , by sea and still be able to germinate. This figure has been questioned based on the extremely small sample size that forms the basis of the paper that makes this claim. Thor Heyerdahl provides an alternative, and much shorter, estimate based on his first-hand experience crossing the Pacific Ocean on the raft ''Kon-Tiki'':
"The nuts we had in baskets on deck remained edible and capable of germinating the whole way to Polynesia. But we had laid about half among the special provisions below deck, with the waves washing around them. Every single one of these was ruined by the sea water. And no coconut can float over the sea faster than a balsa raft moves with the wind behind it."Heyerdahl, Thor. (1950) Kon-Tiki: Across the Pacific by Raft. Mattituck: Amereon House. 240 p.
He also notes that several of the nuts began to germinate by the time they had been ten weeks at sea, precluding an unassisted journey of 100 days or more. Drift models based on wind and ocean currents have shown that coconuts could not have drifted across the Pacific unaided. If they were naturally distributed and had been in the Pacific for a thousand years or so, then we would expect the eastern shore of Australia, with its own islands sheltered by the Great Barrier Reef, to have been thick with coconut palms: the currents were directly into, and down along this coast. However, both James Cook and William Bligh (put adrift after the Mutiny on the Bounty, ''Bounty'' mutiny) found no sign of the nuts along this stretch when he needed water for his crew. Nor were there coconuts on the east side of the African coast until Vasco da Gama, nor in the Caribbean when first visited by Christopher Columbus. They were commonly carried by Spanish ships as a source of fresh water. These provide substantial circumstantial evidence that deliberate Austronesian peoples, Austronesian voyagers were involved in carrying coconuts across the Pacific Ocean and that they could not have dispersed worldwide without human agency. More recently, genomic analysis of cultivated coconut (''C. nucifera'' L.) has shed light on the movement. However, Genetic admixture, admixture, the transfer of genetic material, evidently occurred between the two populations. Given that coconuts are ideally suited for inter-island group ocean dispersal, obviously some natural distribution did take place. However, the locations of the admixture events are limited to Madagascar and coastal east Africa, and exclude the Seychelles. This pattern coincides with the known trade routes of Austronesian sailors. Additionally, a genetically distinct subpopulation of coconut on the Pacific coast of Latin America has undergone a genetic bottleneck resulting from a founder effect; however, its ancestral population is the Pacific coconut from the Philippines. This, together with their use of the South American sweet potato, suggests that Austronesian peoples may have sailed as far east as the Americas. Specimens have been collected from the sea as far north as Norway (but it is not known where they entered the water). In the Hawaiian Islands, the coconut is regarded as a Polynesian introduced species, introduction, first brought to the islands by early Polynesian voyagers from their homelands in the southern islands of Polynesia. They have been found in the Caribbean and the Atlantic coasts of Africa and South America for less than 500 years (the Caribbean native inhabitants do not have a dialect term for them, but use the Portuguese name), but evidence of their presence on the Pacific coast of South America antedates Christopher Columbus's arrival in the Americas. They are now almost ubiquitous between 26° N and 26° S except for the interiors of Africa and South America. The 2014 coral atoll origin hypothesis proposed that the coconut had dispersed in an island hopping fashion using the small, sometimes transient, coral atolls. It noted that by using these small atolls, the species could easily island-hop. Over the course of evolutionary time-scales the shifting atolls would have shortened the paths of colonization, meaning that any one coconut would not have to travel very far to find new land.


Ecology


Pests and diseases

Coconuts are susceptible to the phytoplasma disease, lethal yellowing. One recently selected cultivar, the Maypan coconut palm, 'Maypan', has been bred for resistance to this disease. Yellowing diseases affect plantations in Africa, India, Mexico, the Caribbean and the Pacific Region. The coconut palm is damaged by the larvae of many Lepidoptera (butterfly and moth) species which feed on it, including the African armyworm (''Spodoptera exempta'') and ''Batrachedra'' spp.: ''B. arenosella'', ''B. atriloqua'' (feeds exclusively on ''C. nucifera''), ''B. mathesoni'' (feeds exclusively on ''C. nucifera''), and ''B. nuciferae''. ''Brontispa longissima'' (coconut leaf beetle) feeds on young leaves, and damages both seedlings and mature coconut palms. In 2007, the Philippines imposed a quarantine in Metro Manila and 26 provinces to stop the spread of the pest (organism), pest and protect the Philippine coconut industry managed by some 3.5 million farmers. The fruit may also be damaged by eriophyid coconut mites (''Eriophyes guerreronis''). This mite infests coconut plantations, and is devastating; it can destroy up to 90% of coconut production. The immature seeds are infested and desapped by larvae staying in the portion covered by the perianth of the immature seed; the seeds then drop off or survive deformed. Spraying with wettable sulfur 0.4% or with Neem-based pesticides can give some relief, but is cumbersome and labor-intensive. In Kerala, India, the main coconut pests are the coconut mite, the rhinoceros beetle, the red palm weevil, and the coconut leaf caterpillar. Research into countermeasures to these pests has yielded no results; researchers from the Kerala Agricultural University and the Central Plantation Crop Research Institute, Kasaragode, continue to work on countermeasures. The Krishi Vigyan Kendra, Kannur under Kerala Agricultural University has developed an innovative extension approach called the compact area group approach to combat coconut mites.


Production and cultivation

In 2019, world production of coconuts was 62 million tonnes, led by Indonesia, the Philippines, and India, with 75% combined of the total (table).


Cultivation

Coconut palms are normally cultivated in hot and wet tropical climates. They need year round warmth and moisture to grow well and fruit. Coconut palms are hard to establish in dry climates, and cannot grow there without frequent irrigation; in drought conditions, the new leaves do not open well, and older leaves may become desiccated; fruit also tends to be shed. The extent of cultivation in the tropics is threatening a number of habitats, such as mangroves; an example of such damage to an ecoregion is in the Petenes mangroves of the Yucatán region, Yucatán.


Cultivars

Coconut has a number of commercial and traditional cultivars. They can be sorted mainly into tall cultivars, dwarf cultivars, and hybrid cultivars (hybrids between talls and dwarfs). Some of the dwarf cultivars such as 'Malayan dwarf' have shown some promising resistance to lethal yellowing, while other cultivars such as 'Jamaican tall' are highly affected by the same plant disease. Some cultivars are more drought resistant such as 'West coast tall' (India) while others such as 'Hainan Tall' (China) are more cold tolerant. Other aspects such as seed size, shape and weight, and copra thickness are also important factors in the selection of new cultivars. Some cultivars such as 'Fiji dwarf' form a large bulb at the lower stem and others are cultivated to produce very sweet coconut water with orange-coloured husks (king coconut) used entirely in fruit stalls for drinking (Sri Lanka, India).


Harvesting

In some parts of the world (Thailand and Malaysia), trained Southern pig-tailed macaque, pig-tailed macaques are used to harvest coconuts. Thailand has been raising and training pig-tailed macaques to pick coconuts for around 400 years.What's Funny About The Business Of Monkeys Picking Coconuts?
NPR, October 19, 2015.
Training schools for pig-tailed macaques still exist both in southern Thailand and in the Malaysian state of Kelantan.


Substitutes for cooler climates

In cooler climates (but not less than Hardiness zone, USDA Zone 9), a similar palm, the queen palm (''Syagrus romanzoffiana''), is used in landscaping. Its fruits are similar to the coconut, but smaller. The queen palm was originally classified in the genus ''Cocos'' along with the coconut, but was later reclassified in ''Syagrus (plant), Syagrus''. A recently discovered palm, ''Beccariophoenix alfredii'' from Madagascar, is nearly identical to the coconut, more so than the queen palm and can also be grown in slightly cooler climates than the coconut palm. Coconuts can only be grown in temperatures above and need a daily temperature above to produce fruit.


Production by country


Indonesia

Indonesia is the world's largest producer of coconuts, with gross production of 15 million tonnes. A sprouting coconut seed is the logo for Gerakan Pramuka Indonesia, the Indonesian scouting organization.


Philippines

The Philippines is the world's second-largest producer of coconuts. It was the world's largest producer for decades until a decline in production due to aging trees as well as typhoon devastation. Indonesia overtook it in 2010. It is still the largest producer of coconut oil and copra, accounting for 64% of the global production. The production of coconuts plays an important role in the Economy of the Philippines, economy, with 25% of cultivated land (around 3.56 million hectares) used for coconut plantations and approximately 25 to 33% of the population reliant on coconuts for their livelihood. Two important coconut products were first developed in the Philippines, macapuno and ''nata de coco''. Macapuno is a coconut variety with a jelly-like coconut meat. Its meat is sweetened, cut into strands, and sold in glass jars as coconut strings, sometimes labeled as "gelatinous mutant coconut". ''Nata de coco'', also called coconut gel, is another jelly-like coconut product made from fermented coconut water.


India

Traditional areas of coconut cultivation in India are the states of Kerala, Tamil Nadu, Karnataka, Puducherry (union territory), Puducherry, Andhra Pradesh, Goa, Maharashtra, Odisha, West Bengal and, Gujarat and the islands of Lakshadweep and Andaman and Nicobar. As per 2014–15 statistics from Coconut Development Board of Government of India, four southern states combined account for almost 90% of the total production in the country: Tamil Nadu (33.84%), Karnataka (25.15%), Kerala (23.96%), and Andhra Pradesh (7.16%). Other states, such as Goa, Maharashtra, Odisha, West Bengal, and those in the northeast (Tripura and Assam) account for the remaining productions. Though Kerala has the largest number of coconut trees, in terms of production per hectare, Tamil Nadu leads all other states. In Tamil Nadu, Coimbatore and Tirupur regions top the production list. In Goa, the coconut tree has been reclassified by the government as a palm (like a grass), enabling farmers and real estate developers to clear land with fewer restrictions. With this, it will no more be considered as a tree and no permission will be required by the forest department before cutting a coconut tree.


Middle East

The main coconut-producing area in the Middle East is the Dhofar region of Oman, but they can be grown all along the Persian Gulf, Arabian Sea, and Red Sea coasts, because these seas are tropical and provide enough humidity (through seawater evaporation) for coconut trees to grow. The young coconut plants need to be nursed and irrigated with drip pipes until they are old enough (stem bulb development) to be irrigated with brackish water or seawater alone, after which they can be replanted on the beaches. In particular, the area around Salalah maintains large coconut plantations similar to those found across the Arabian Sea in Kerala. The reasons why coconut are cultivated only in Yemen's Al Mahrah and Hadramaut governorates and in the Sultanate of Oman, but not in other suitable areas in the Arabian Peninsula, may originate from the fact that Oman and Hadramaut had long dhow trade relations with Burma, Malaysia, Indonesia, East Africa, and Zanzibar, as well as southern India and China. Omani people needed the coir rope from the coconut fiber to stitch together their traditional seagoing dhow vessels in which nails were never used. The knowhow of coconut cultivation and necessary Erosion control, soil fixation and irrigation may have found its way into Omani, Hadrami and Al-Mahra culture by people who returned from those overseas areas. The coconut cultivars grown in Oman are generally of the drought-resistant Indian 'West Coast tall' variety. Unlike the UAE, which grows mostly non-native dwarf or hybrid coconut cultivars imported from Florida for ornamental purposes, the slender, tall Omani coconut cultivars are relatively well-adapted to the Middle East's hot dry seasons, but need longer to reach maturity. The Middle East's hot, dry climate favors the development of coconut mites, which cause immature seed dropping and may cause brownish-gray discoloration on the coconut's outer green fiber. The ancient coconut groves of Dhofar were mentioned by the medieval Moroccan traveller Ibn Battuta in his writings, known as ''Al Rihla''. The annual rainy season known locally as ''khareef'' or monsoon makes coconut cultivation easy on the Arabian east coast. Coconut trees also are increasingly grown for decorative purposes along the coasts of the United Arab Emirates and Saudi Arabia with the help of irrigation. The UAE has, however, imposed strict laws on mature coconut tree imports from other countries to reduce the spread of pest (organism), pests to other native palm trees, as the mixing of date and coconut trees poses a risk of cross-species palm pests, such as rhinoceros beetles and red palm weevils. The artificial landscaping may have been the cause for lethal yellowing, a viral coconut palm disease that leads to the death of the tree. It is spread by host insects, that thrive on heavy turf grasses. Therefore, heavy turf grass environments (beach resorts and golf courses) also pose a major threat to local coconut trees. Traditionally, dessert banana plants and local wild beach flora such as ''Scaevola taccada'' and ''Ipomoea pes-caprae'' were used as humidity-supplying green undergrowth for coconut trees, mixed with sea almond and sea hibiscus. Due to growing sedentary lifestyles and heavy-handed landscaping, a decline in these traditional farming and soil-fixing techniques has occurred.


Sri Lanka

Sri Lanka is the world's fourth-largest producer of coconuts and is the second-largest producer of coconut oil and copra, accounting for 15% of the global production. The production of coconuts is the main source of Economy of Sri Lanka, Sri Lanka economy, with 12% of cultivated land and 409,244 hectares used for coconut growing (2017). Sri Lanka established its Coconut Development Authority and Coconut Cultivation Board and Coconut Research Institute in the early British Ceylon period.


United States

In the United States, coconut palms can be grown and reproduced outdoors without irrigation in Hawaii, southern and central Florida, and the territories of Puerto Rico, Guam, American Samoa, the U.S. Virgin Islands, and the Commonwealth of the Northern Mariana Islands, Northern Mariana Islands. In Florida, wild populations of coconut palms extend up the East Coast from Key West to Jupiter Inlet, and up the West Coast from Marco Island to Sarasota. Many of the smallest coral islands in the Florida Keys are known to have abundant coconut palms sprouting from coconuts that have drifted or been deposited by ocean currents. Coconut palms are cultivated north of south Florida to roughly Cocoa Beach on the East Coast and Clearwater, Florida, Clearwater on the West Coast.


Australia

Coconuts are commonly grown around the northern coast of Australia, and in some warmer parts of New South Wales. However they are mainly present as decoration, and the Australian coconut industry is small; Australia is a net importer of coconut products. Australian cities put much effort into de-fruiting decorative coconut trees to ensure that the mature coconuts do not fall and injure people.


Uses

The coconut palm is grown throughout the
tropics The tropics are the region of Earth surrounding the Equator. They are delimited in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S; these latitudes correspond to the ...
for decoration, as well as for its many culinary and nonculinary uses; virtually every part of the coconut palm can be used by humans in some manner and has significant economic value. Coconuts' versatility is sometimes noted in its naming. In Sanskrit, it is ''kalpa vriksha'' ("the tree which provides all the necessities of life"). In the Malay language, it is ''pokok seribu guna'' ("the tree of a thousand uses"). In the Philippines, the coconut is commonly called the "tree of life". It is one of the most useful trees in the world.


Culinary uses

The coconut flesh can be cut into strips, salted and baked to make bacon-like fixings.


Nutrition

A reference serving of raw coconut flesh supplies of food energy and a high amount of total fat (33 grams), especially saturated fat (89% of total fat), along with a moderate quantity of carbohydrates (15 grams), and protein (3 grams). Micronutrients in significant content (more than 10% of the Daily Value) include the dietary minerals, manganese, copper, iron, phosphorus, selenium, and zinc (table). The various parts of the coconut have a number of culinary uses.


Flaked coconut

The white, fleshy part of the seed, the coconut meat, is used fresh or dried in cooking, especially in confections and desserts such as macaroons and buko pie. Dried coconut is also used as the filling for many chocolate bars. Some dried coconut is purely coconut, but others are manufactured with other ingredients, such as sugar, propylene glycol, salt, and sodium metabisulfite. Fresh shredded or flaked coconut is also used as a Garnish (food), garnish various dishes, as in ''klepon'' and ''puto bumbóng''.


Macapuno

A special cultivar of coconut known as macapuno has a jelly-like coconut meat. It was first developed for commercial cultivation in the Philippines and is used widely in Philippine cuisine for desserts, drinks, and pastries. It is also popular in Indonesia (where it is known as ''kopyor'') for making beverages.


Coconut milk

Coconut milk, not to be confused with coconut water, is obtained by pressing the grated coconut meat, usually with hot water added which extracts the coconut oil, proteins, and aromatic compounds. It is used for cooking various dishes. Coconut milk contains 5% to 20% fat, while coconut cream contains around 20% to 50% fat. Most of which (89%) is saturated fat, with lauric acid as a major fatty acid. Coconut milk can be diluted to create coconut milk beverages. These have much lower fat content and are suitable as milk substitutes. The milk can be used to produce virgin coconut oil by controlled heating and removal of the oil fraction. Coconut milk powder, a protein-rich powder can be processed from coconut milk following centrifugation, separation process, separation, and spray drying. Coconut milk and coconut cream extracted from grated coconut is frequently added to various dessert and savory dishes, as well as in curries and stews. It can also be diluted into a beverage. Various other products made from thickened coconut milk with sugar and/or eggs like coconut jam and coconut custard are also widespread in Southeast Asia. In the Philippines, sweetened reduced coconut milk is marketed as coconut syrup and is used for various desserts. Coconut oil extracted from coconut milk or copra is also used for frying, cooking, and making margarine, among other uses.


Coconut water

Coconut water serves as a suspension for the endosperm of the coconut during its Cell cycle#Mitotic phase (chromosome separation), nuclear phase of development. Later, the endosperm matures and deposits onto the coconut rind during the cellular phase. It is consumed throughout the humid tropics, and has been introduced into the retail market as a processed sports drink. Mature fruits have significantly less liquid than young, immature coconuts, barring spoilage. Coconut water can be fermented to produce coconut vinegar. Per 100-gram serving, coconut water contains 19 calories and no significant content of essential nutrients. Coconut water can be drunk fresh or used in cooking as in ''binakol''. It can also be fermented to produce a jelly-like dessert known as ''nata de coco''.


Coconut flour

Coconut flour has also been developed for use in baking, to combat malnutrition.


Heart of palm

Apical meristem, Apical buds of adult plants are edible, and are known as "palm cabbage" or heart of palm. They are considered a rare delicacy, as harvesting the buds kills the palms. Hearts of palm are eaten in salads, sometimes called "millionaire's salad".


Sprouted coconut

Newly germinated coconuts contain an edible fluff of marshmallow-like consistency called sprouted coconut or coconut sprout, produced as the endosperm nourishes the Plant embryogenesis, developing embryo. It is a haustorium, a spongy absorbent tissue formed from the distal portion of embryo during coconut germination, facilitates absorption of nutrients for the growing shoot and root.


Toddy and sap

The sap derived from incising the flower clusters of the coconut is drunk as ''neera'', also known as toddy or ''tubâ'' (Philippines), ''tuak'' (Indonesia and Malaysia) or ''karewe'' (fresh and not fermented, collected twice a day, for breakfast and dinner) in Kiribati. When left to ferment on its own, it becomes palm wine. Palm wine is distilled to produce ''arrack''. In the Philippines, this alcoholic drink is called ''Arrack#Coconut arrack, lambanog'' or "coconut vodka". The sap can be reduced by boiling to create a sweet syrup or candy such as ''te kamamai'' in Kiribati or ''dhiyaa hakuru'' and ''addu bondi'' in the Maldives. It can be reduced further to yield coconut sugar also referred to as palm sugar or jaggery. A young, well-maintained tree can produce around of toddy per year, while a 40-year-old tree may yield around . Coconut sap, usually extracted from cut inflorescence stalks is sweet when fresh and can be drunk as is like in ''tuba fresca'' of Mexico. They can also be processed to extract palm sugar. The sap when fermented can also be made into coconut vinegar or various palm wines (which can be further distilled to make arrack).


Coconut oil

Coconut oil is commonly used in cooking, especially for frying. It can be used in liquid form as would other vegetable oils, or in solid form similar to butter or lard. Long-term consumption of coconut oil may have negative health effects similar to those from consuming other sources of saturated fats, including butter, beef fat, and palm oil. Its chronic consumption may increase the risk of cardiovascular diseases by raising total blood cholesterol levels through elevated blood levels of low-density lipoprotein, LDL cholesterol and lauric acid.


Coconut butter

Coconut butter is often used to describe solidified coconut oil, but has also been adopted as an alternate name for creamed coconut, a specialty product made of coconut milk solids or puréed coconut meat and oil.


Coir

Coir (the fiber from the husk of the coconut) is used in ropes, mats, mat, doormats, brushes, and sacks, as caulking for boats, and as stuffing fiber for mattresses. It is used in horticulture in potting compost, especially in orchid mix. The coir is used to make brooms in Cambodia.


Copra

Copra is the dried meat of the seed and after processing produces coconut oil and coconut meal. Coconut oil, aside from being used in cooking as an ingredient and for frying, is used in soaps, cosmetics, hair oil, and massage oil. Coconut oil is also a main ingredient in Ayurveda, Ayurvedic oils. In Vanuatu, coconut palms for copra production are generally spaced apart, allowing a tree density of .


Husks and shells

The husk and shells can be used for fuel and are a source of
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charcoal
. Activated carbon manufactured from coconut shell is considered extremely effective for the removal of impurities. The coconut's obscure origin in foreign lands led to the notion of using cups made from the shell to neutralise poisoned drinks. The cups were frequently engraved and decorated with precious metals. A dried half coconut shell with husk can be used to buff floors. It is known as a ''bunot'' in the Philippines and simply a "coconut brush" in Jamaica. The fresh husk of a brown coconut may serve as a dish sponge or body sponge. A ''coco chocolatero'' was a cup used to serve small quantities of beverages (such as chocolate drinks) between the 17th and 19th centuries in countries such as Mexico, Guatemala, and Venezuela. In Asia, coconut shells are also used as bowls and in the manufacture of various handicrafts, including buttons carved from dried shell. Coconut buttons are often used for Hawaiian aloha shirts. ''Tempurung'', as the shell is called in the Malay language, can be used as a soup bowl and—if fixed with a handle—a ladle. In Thailand, the coconut husk is used as a potting medium to produce healthy forest tree saplings. The process of husk extraction from the coir bypasses the retting process, using a custom-built coconut husk extractor designed by ASEAN–Canada Forest Tree Seed Centre in 1986. Fresh husks contain more tannin than old husks. Tannin produces negative effects on sapling growth. In parts of South India, the shell and husk are burned for smoke to repel mosquitoes. Half coconut shells are used in theatre Foley sound effects work, struck together to create the sound effect of a horse's hoofbeats. Dried half shells are used as the bodies of musical instruments, including the Chinese ''yehu'' and ''banhu'', along with the Vietnamese ''đàn gáo'' and Arabo-Turkic ''rebab''. In the Philippines, dried half shells are also used as a music instrument in a folk dance called ''maglalatik''. The shell, freed from the husk, and heated on warm ashes, exudes an oily material that is used to soothe dental pains in traditional medicine of Cambodia. In World War II, coastwatchers, coastwatcher scout Biuku Gasa was the first of two from the Solomon Islands to reach the shipwrecked and wounded crew of Motor Torpedo Boat PT-109, Motor Torpedo Boat ''PT-109'' commanded by future U.S. president John F. Kennedy. Gasa suggested, for lack of paper, delivering by dugout canoe a message inscribed on a husked coconut shell, reading “Nauru Isl commander / native knows posit / he can pilot / 11 alive need small boat / Kennedy.” This coconut was later kept on the president's desk, and is now in the John F. Kennedy Library.


Leaves

The stiff midribs of coconut leaves are used for making brooms in India, Indonesia (''sapu lidi''), Malaysia, the Maldives, and the Philippines (''walis tingting''). The green of the leaves (lamina) is stripped away, leaving the veins (long, thin, woodlike strips) which are tied together to form a broom or brush. A long handle made from some other wood may be inserted into the base of the bundle and used as a two-handed broom. The leaves also provide material for baskets that can draw well water and for roofing thatch; they can be woven into mats, cooking skewers, and kindling arrows as well. Leaves are also woven into small piuches that are filled with rice and cooked to make ''pusô'' and ''ketupat''. Dried coconut leaves can be burned to ash, which can be harvested for lime (mineral), lime. In India, the woven coconut leaves are used to build wedding marquee (tent), marquees, especially in the states of Kerala, Karnataka, and Tamil Nadu. The leaves are used for thatching houses, or for decorating climbing frames and meeting rooms in Cambodia, where the plant is known as ''dôô:ng''.


Timber

Coconut trunks are used for building small bridges and huts; they are preferred for their straightness, strength, and salt resistance. In Kerala, coconut trunks are used for house construction. Coconut timber comes from the trunk, and is increasingly being used as an ecologically sound substitute for endangered hardwoods. It has applications in furniture and specialized construction, as notably demonstrated in Manila's Coconut Palace. Hawaiians hollowed the trunk to form drums, containers, or small canoes. The "branches" (leaf petiole (botany), petioles) are strong and flexible enough to make a switch (rod), switch. The use of coconut branches in corporal punishment was revived in the Gilbertese community on Choiseul in the Solomon Islands in 2005.


Roots

The roots are used as a dye, a mouthwash, and a folk medicine for diarrhea and dysentery. A frayed piece of root can also be used as a toothbrush. In Cambodia, the roots are used in traditional medicine as a treatment for dysentery.


Other uses

The leftover fiber from coconut oil and coconut milk production, coconut meal, is used as livestock feed. The dried calyx (botany), calyx is used as fuel in wood-fired stoves. Coconut water is traditionally used as a growth supplement in plant tissue culture and micropropagation. The smell of coconuts comes from the δ-Decalactone, 6-pentyloxan-2-one molecule, known as δ-decalactone in the food and fragrance industries.


Tool and shelter for animals

Researchers from the Melbourne Museum in Australia observed the octopus species ''Amphioctopus marginatus'' tool use by animals, use tools, specifically coconut shells, for defense and shelter. The discovery of this behavior was observed in Bali and North Sulawesi in Indonesia between 1998 and 2008. ''Amphioctopus marginatus'' is the first invertebrate known to be able to use tools. A coconut can be hollowed out and used as a home for a rodent or small birds. Halved, drained coconuts can also be hung up as bird feeders, and after the flesh has gone, can be filled with fat in winter to attract tit (bird), tits.


Allergies


Food allergies

Coconut oil is increasingly used in the food industry. Proteins from coconut may cause food allergy, including anaphylaxis. In the United States, the U.S. Food and Drug Administration declared that coconut must be disclosed as an ingredient on package labels as a "tree nut" with potential allergenicity.


Topical allergies

Cocamidopropyl betaine (CAPB) is a surfactant manufactured from coconut oil that is increasingly used as an ingredient in personal hygiene products and cosmetics, such as shampoos, liquid soaps, cleansers and antiseptics, among others. CAPB may cause mild skin irritation, but allergic reactions to CAPB are rare and probably related to impurities rendered during the manufacturing process (which include amidoamine and dimethylaminopropylamine) rather than CAPB itself.


In culture

The coconut was a critical food item for the people of Polynesia, and the Polynesians brought it with them as they spread to new islands. In the Ilocos region of the northern Philippines, the Ilocano people fill two halved coconut shells with ''diket'' (cooked sweet rice), and place ''liningta nga itlog'' (halved boiled egg) on top of it. This ritual, known as ''niniyogan'', is an offering made to the deceased and one's ancestors. This accompanies the ''palagip'' (prayer to the dead). A coconut ( sa, narikela) is an essential element of rituals in Hindu tradition. Often it is decorated with bright metal foils and other symbols of auspiciousness. It is offered during worship to a Hindu god or goddess. Narali Purnima is celebrated on a full moon day which usually signifies the end of monsoon season in India. The word ''Narali'' is derived from ''naral'' implying "coconut" in Marathi people, Marathi. Fishermen give an offering of coconut to the sea to celebrate the beginning of a new fishing season. Irrespective of their religious affiliations, fishermen of India often offer it to the rivers and seas in the hopes of having bountiful catches. Hindus often initiate the beginning of any new activity by breaking a coconut to ensure the blessings of the gods and successful completion of the activity. The Hindu goddess of well-being and wealth, Lakshmi, is often shown holding a coconut. In the foothills of the temple town of Palani, before going to worship Murugan for the Ganesha, coconuts are broken at a place marked for the purpose. Every day, thousands of coconuts are broken, and some devotees break as many as 108 coconuts at a time as per the prayer. They are also used in Hindu weddings as a symbol of prosperity. The flowers are used sometimes in wedding ceremonies in Cambodia. The Zulu Social Aid and Pleasure Club of New Orleans traditionally throws hand-decorated coconuts, one of the most valuable Mardi Gras souvenirs, to parade revelers. The tradition began in the 1910s, and has continued since. In 1987, a "coconut law" was signed by Governor Edwin Edwards exempting from insurance liability any decorated coconut "handed" from a Zulu float. The coconut is also used as a target and prize in the traditional British fairground game coconut shy. The player buys some small balls which are then thrown as hard as possible at coconuts balanced on sticks. The aim is to knock a coconut off the stand and win it. It was the main food of adherents of the now discontinued Vietnamese religion Coconut Religion, Đạo Dừa.


Myths and legends

Some South Asian, Southeast Asian, and Pacific Ocean cultures have origin myths in which the coconut plays the main role. In the Hainuwele myth from Maluku Islands, Maluku, a girl emerges from the blossom of a coconut tree. In Maldivian folklore, one of the main myths of origin reflects the dependence of the Maldivians on the coconut tree.Xavier Romero-Frias, Romero-Frias, Xavier (2012) ''Folk tales of the Maldives''
NIAS Press
, ,
In the story of Sina and the Eel, the origin of the coconut is related as the beautiful woman Sina burying an eel, which eventually became the first coconut. According to urban legend, death by coconut, more deaths are caused by falling coconuts than by sharks annually.


See also

* Domesticated plants and animals of Austronesia * Central Plantation Crops Research Institute * Coconut production in Kerala * Coir Board of India * List of coconut dishes * List of dishes made using coconut milk * Ravanahatha a musical instrument sometimes made of coconuts * ''Voanioala gerardii'' forest coconut, the closest relative of the modern coconut


References


Further reading

* Adkins S.W., M. Foale and Y.M.S. Samosir (eds.) (2006)
Coconut revival – new possibilities for the ‘tree of life’
Proceedings of the International Coconut Forum held in Cairns, Australia, November 22–24, 2005. ACIAR Proceedings No. 125. * * Frison, E.A.; Putter, C.A.J.; Diekmann, M. (eds.). (1993)

. * International Plant Genetic Resources Institute (IPGRI). (1995)

. * Mathur, P.N.; Muralidharan, K.; Parthasarathy, V.A.; Batugal, P.; Bonnot, F. (2008)

. * Salunkhe, D.K., J.K. Chavan, R.N. Adsule, and S.S. Kadam. (1992). ''World Oilseeds – Chemistry, Technology, and Utilization''. Springer. .


External links

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