Cladonia Amaurocraea
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''Cladonia amaurocraea'', commonly known as the quill lichen or the quill cup lichen, is a species of
fruticose A fruticose lichen is a form of lichen fungi that is characterized by a coral-like shrubby or bushy growth structure. It is formed from a symbiotic relationship of a photobiont such as green algae or less commonly cyanobacteria and one, two or ...
,
cup lichen A cup is an open-top used to hold hot or cold liquids for pouring or drinking; while mainly used for drinking, it also can be used to store solids for pouring (e.g., sugar, flour, grains, salt). Cups may be made of glass, metal, china, clay, ...
in the
Cladoniaceae The Cladoniaceae are a family of lichenized fungi in the order Lecanorales. It is one of the largest families of lichen-forming fungi, with about 560 species distributed amongst 17 genera. The reindeer moss and cup lichens (''Cladonia'') belong t ...
family.


Taxonomy

The lichen was first described scientifically as ''Capitularia amaurocraea'' in 1810. It was transferred to the genus ''
Cladonia ''Cladonia'' is a genus of moss-like lichens in the family Cladoniaceae. They are the primary food source for reindeer/caribou. ''Cladonia'' species are of economic importance to reindeer-herders, such as the Sami in Scandinavia or the Nenets ...
'' in 1887. It is commonly known as quill lichen.


Description

The
thallus Thallus (plural: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. Many of these organisms wer ...
of ''Cladonia amaurocraea'' comprises tall (15–100 mm high) and slender
podetia A podetium (plural: podetia) is the upright secondary thallus in ''Cladonia'' lichens. It is a hollow stalk extending from the primary thallus. Podetia can be pointed stalks, club like, cupped, or branched in shape and may or may not contain the a ...
that are irregularly or dichotomously branched. These podetia have a smooth, yellowish-green surface that is often mottled with patches of green and white. They either form a pointy tip, or a narrow cup that is either closed or has a narrow opening. The
cortex Cortex or cortical may refer to: Biology * Cortex (anatomy), the outermost layer of an organ ** Cerebral cortex, the outer layer of the vertebrate cerebrum, part of which is the ''forebrain'' *** Motor cortex, the regions of the cerebral cortex i ...
contains
usnic acid Usnic acid is a naturally occurring dibenzofuran derivative found in several lichen species with the formula C18H16O7. It was first isolated by German scientist W. Knop in 1844 and first synthesized between 1933-1937 by Curd and Robertson. Usnic a ...
, while the
medulla Medulla or Medullary may refer to: Science * Medulla oblongata, a part of the brain stem * Renal medulla, a part of the kidney * Adrenal medulla, a part of the adrenal gland * Medulla of ovary, a stroma in the center of the ovary * Medulla of t ...
has
barbatic acid Barbatic acid is an organic compound that is made by some lichens. It is in the structural class known as depsides. It is particularly common in the genera ''Usnea'' (the beard lichens) and ''Cladonia''. History The compound was first isolated ...
.


Habitat and distribution

''Cladonia amaurocraea'' is found in
boreal forest Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, ...
s, where it typically grows on talus deposits between boulders and on rocky ground. In North America, it is widespread throughout Canada and Alaska.


See also

* List of ''Cladonia'' species


References

amaurocraea Lichen species Lichens described in 1810 Lichens of Europe Lichens of North America Taxa named by Heinrich Gustav Flörke {{Lecanorales-stub