Charles Roy Henderson
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Charles Roy Henderson ( – ) was an American statistician and a pioneer in
animal breeding Animal breeding is a branch of animal science that addresses the evaluation (using best linear unbiased prediction and other methods) of the genetic value (estimated breeding value, EBV) of livestock. Selecting for breeding animals with superior E ...
— the application of quantitative methods for the genetic evaluation of domestic livestock. This is critically important because it allows farmers and geneticists to predict whether a crop or animal will have a desired trait, and to what extent the trait will be expressed. He developed
mixed model A mixed model, mixed-effects model or mixed error-component model is a statistical model containing both fixed effects and random effects. These models are useful in a wide variety of disciplines in the physical, biological and social sciences. ...
equations to obtain
best linear unbiased prediction In statistics, best linear unbiased prediction (BLUP) is used in linear mixed models for the estimation of random effects. BLUP was derived by Charles Roy Henderson in 1950 but the term "best linear unbiased predictor" (or "prediction") seems not ...
s of breeding values and, in general, any random effect. He invented three methods for the estimation of
variance components In statistics, a random effects model, also called a variance components model, is a statistical model where the model parameters are random variables. It is a kind of hierarchical linear model, which assumes that the data being analysed are ...
in unbalanced settings of mixed models, and invented a method for constructing the inverse of Wright's numerator relationship matrix based on a simple list of pedigree information. He, with his Ph.D. student Shayle R. Searle, greatly extended the use of matrix notation in statistics. His methods are widely used by the domestic livestock industry throughout the world and are a cornerstone of linear model theory. Henderson obtained his
B.Sc. A Bachelor of Science (BS, BSc, SB, or ScB; from the Latin ') is a bachelor's degree awarded for programs that generally last three to five years. The first university to admit a student to the degree of Bachelor of Science was the University of ...
,
M.Sc. A Master of Science ( la, Magisterii Scientiae; abbreviated MS, M.S., MSc, M.Sc., SM, S.M., ScM or Sc.M.) is a master's degree in the field of science awarded by universities in many countries or a person holding such a degree. In contrast to ...
(nutrition) and
Ph.D. A Doctor of Philosophy (PhD, Ph.D., or DPhil; Latin: or ') is the most common degree at the highest academic level awarded following a course of study. PhDs are awarded for programs across the whole breadth of academic fields. Because it is ...
(breeding) degrees at
Iowa State University Iowa State University of Science and Technology (Iowa State University, Iowa State, or ISU) is a public land-grant research university in Ames, Iowa. Founded in 1858 as the Iowa Agricultural College and Model Farm, Iowa State became one of the ...
, where he was a student of Professor L. N. Hazel. Henderson joined the faculty of the Department of Animal Science at
Cornell University Cornell University is a private statutory land-grant research university based in Ithaca, New York. It is a member of the Ivy League. Founded in 1865 by Ezra Cornell and Andrew Dickson White, Cornell was founded with the intention to tea ...
in 1948, and headed the Animal Breeding division for nearly 30 years until he retired in 1976. After retiring from Cornell, he was a visiting professor at the
University of Guelph , mottoeng = "to learn the reasons of realities" , established = May 8, 1964 ()As constituents: OAC: (1874) Macdonald Institute: (1903) OVC: (1922) , type = Public university , chancellor ...
and
University of Illinois The University of Illinois Urbana-Champaign (U of I, Illinois, University of Illinois, or UIUC) is a public land-grant research university in Illinois in the twin cities of Champaign and Urbana. It is the flagship institution of the Univer ...
until his death. He completed his book in 1984 at the
University of Guelph , mottoeng = "to learn the reasons of realities" , established = May 8, 1964 ()As constituents: OAC: (1874) Macdonald Institute: (1903) OVC: (1922) , type = Public university , chancellor ...
.


Honors and awards

* 1955, Senior Fulbright Research Scholar (New Zealand) * 1964, Borden Award,
American Dairy Science Association The American Dairy Science Association (ADSA) is a non-profit professional organization for the advancement of dairy science. ADSA is headquartered in Champaign, Illinois. Consisting of 4500 members, ADSA is involved in research, education, and in ...
* 1964, Animal Breeding and Genetics Award, American Society of Animal Science * 1969, Fellow,
American Statistical Association The American Statistical Association (ASA) is the main professional organization for statisticians and related professionals in the United States. It was founded in Boston, Massachusetts on November 27, 1839, and is the second oldest continuousl ...
* 1971, Morrison Award, American Society of Animal Science * 1977, National Association of Animal Breeders Award, American Dairy Science Association * 1980, Member of the Massey University Wellington * 1981, Visiting professor at the
University of California, Davis The University of California, Davis (UC Davis, UCD, or Davis) is a public land-grant research university near Davis, California. Named a Public Ivy, it is the northernmost of the ten campuses of the University of California system. The inst ...
* 1981, Hermann-von-Nathusius-Medaille in Gold of the German Society for Animal Production (DGfZ) List of people honored with the Herman-von-Nathusius-Medaille
/ref> * 1981, Fellow, American Society of Animal Science * 1982, Jay L. Lush Animal Breeding and Genetics Award, American Dairy Science Association * 1984, Henry A. Wallace Award for Service to Agriculture, Iowa State University * 1985, Elected to the
United States National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the Nati ...


Selected bibliography

;1990s * * ;1980s * * * * * * * * * * * * * * * * * * ;1970s * * * * * * * * * * * * * * * * * * * * ;1960s * * * * * * * * * * * * ;1950s * * *


References

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Footnotes


External links

* * {{DEFAULTSORT:Henderson, Charles Roy 1911 births 1989 deaths Biostatisticians American geneticists Iowa State University alumni Cornell University faculty Fellows of the American Statistical Association Members of the United States National Academy of Sciences Deaths from pulmonary embolism 20th-century American mathematicians Fulbright alumni