Chapalmalania
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''Chapalmalania'' is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
procyonid Procyonidae is a New World family of the order Carnivora. It comprises the raccoons, ringtails, cacomistles, coatis, kinkajous, olingos, and olinguitos. Procyonids inhabit a wide range of environments and are generally omnivorous. Characteri ...
from the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Chapadmalalan The Chapadmalalan age is a period of geologic time (4.0–3.0 Ma) within the Pliocene epoch of the Neogene used more specifically with South American Land Mammal Ages. It follows the Montehermosan and precedes the Uquian age. Fossil content F ...
to
Uquian The Uquian age is a period of geologic time (3.0–1.5 Ma) within the Pliocene and Pleistocene epochs of the Neogene used more specifically with South American Land Mammal Ages. It follows the Montehermosan and precedes the Ensenadan The Ensenad ...
) of
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
and
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
(
Ware Formation The Ware Formation ( es, Formación Ware) is a fossiliferous geological formation of the Cocinetas Basin in the northernmost department of La Guajira. The formation consists of fine lithic to quartzitic sandstones, mudstones, pebbly conglomerate ...
,
Cocinetas Basin The Cocinetas Basin ( es, Cuenca Cocinetas) is a small sedimentary basin of approximately in northeasternmost Colombia. The onshore pull-apart basin is located in the department of La Guajira at the border with Zulia, Venezuela. The basin is bou ...
,
La Guajira La Guajira () is a department of Colombia. It occupies most of the Guajira Peninsula in the northeast region of the country, on the Caribbean Sea and bordering Venezuela, at the northernmost tip of South America. The capital city of the departm ...
).


Description

Though related to
raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the common raccoon to distinguish it from other species, is a mammal native to North America. It is the largest of the procyonid family, having a body length of , and a body weight of ...
s and
coati Coatis, also known as coatimundis (), are members of the family Procyonidae in the genera ''Nasua'' and ''Nasuella''. They are diurnal mammals native to South America, Central America, Mexico, and the southwestern United States. The name ...
s, ''Chapalmalania'' was a large creature, reaching about in body length, with a short tail. ''Chapalmalania'' is estimated to have weighed approximately , comparable in size to an
American black bear The American black bear (''Ursus americanus''), also called simply a black bear or sometimes a baribal, is a medium-sized bear endemic to North America. It is the continent's smallest and most widely distributed bear species. American black bear ...
(''Ursus americanus''). Due to its size, its remains were initially identified as those of a
bear Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the Nor ...
. It evolved from the "dog-coati" ''
Cyonasua ''Cyonasua'' (meaning "dog-coati" in Greek) is an extinct genus of procyonid from the Late Miocene to Middle Pleistocene of South America. Fossils of ''Cyonasua'' have been found in Argentina ( Ituzaingó, Epecuén, Huayquerías, Monte Hermoso, ...
'', which probably island-hopped from
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
during the late
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
(7.5 million years ago), as perhaps the earliest southward mammalian migrants of the
Great American Interchange The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which lan ...
. When the
Isthmus of Panama The Isthmus of Panama ( es, Istmo de Panamá), also historically known as the Isthmus of Darien (), is the narrow strip of land that lies between the Caribbean Sea and the Pacific Ocean, linking North and South America. It contains the country ...
rose from the sea to allow further invasions by other
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
n species, ''Chapalmalania'' was unable to compete and its lineage became extinct. ''Chapalmalania'' is thought to have had an omnivorous diet similar to modern bears based on dental morphology. Bite marks attributable to ''Chapalmalania'' have been found on a
glyptodont Glyptodonts are an extinct subfamily of large, heavily armoured armadillos. They arose in South America around 48 million years ago and spread to southern North America after the continents became connected several million years ago. The best-kn ...
carcass and have been interpreted as scavenging behavior, suggesting that ''Chapalmalania'' also fed on
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
of large mammals on at least some occasions.


References


Further reading

*Barry Cox, Colin Harrison, R.J.G. Savage, and Brian Gardiner. (1999): The Simon & Schuster Encyclopedia of Dinosaurs and Prehistoric Creatures: A Visual Who's Who of Prehistoric Life.
Simon & Schuster Simon & Schuster () is an American publishing company and a subsidiary of Paramount Global. It was founded in New York City on January 2, 1924 by Richard L. Simon and M. Lincoln Schuster. As of 2016, Simon & Schuster was the third largest publ ...
. *David Norman. (2001): The Big Book Of Dinosaurs. page 13, Walcome books.


External links

* Prehistoric procyonids Prehistoric carnivoran genera Pliocene carnivorans Piacenzian extinctions Pliocene mammals of South America Uquian Neogene Argentina Fossils of Argentina Neogene Colombia Fossils of Colombia Fossil taxa described in 1908 Taxa named by Florentino Ameghino {{paleo-carnivora-stub