Calycanthaceae
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The Calycanthaceae (sweetshrubs or spicebushes) are a small
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s in the order
Laurales The Laurales are an order of flowering plants. They are magnoliids, related to the Magnoliales. The order includes about 2500-2800 species from 85-90 genera, which comprise seven families of trees and shrubs. Most of the species are tropical an ...
. The family contains three genera and only 10 known species , restricted to warm temperate and tropical regions: * ''
Calycanthus ''Calycanthus'', called sweetshrub, is a genus of flowering plants in the family Calycanthaceae. The genus includes two to four species depending on taxonomic interpretation; three are accepted by most 21st century sources. Description ''Calyca ...
'' (three species; western and southeastern
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
and one species in eastern
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
) * ''
Chimonanthus ''Chimonanthus'' is a genus of flowering plants in the family Calycanthaceae, native to China, but is also cultivated elsewhere in Asia, including Iran. The genus includes three to six species depending on taxonomic interpretation; six are acce ...
'' (six species; eastern
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
) * ''
Idiospermum ''Idiospermum'' is a genus containing a single species of tree, ''Idiospermum australiense'', found in Australian tropical rainforests. The species represents one of the earliest known lineages of flowering plants, dating back as far as 120 milli ...
'' (one species;
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
,
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
) They are aromatic,
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s growing to 2–4 m tall, except for ''Idiospermum'', which is a large
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s are white to red, with spirally arranged
tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s. DNA-based phylogenies indicate the Northern Hemisphere ''Calycanthus'' and ''Chimonanthus'' diverged from each other in the mid-Miocene, while the Australian ''Idiospermum'' had already diverged by the Upper Cretaceous and likely represents a remnant of a former
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
n distribution of Calycanthaceae that included South America, as indicated by the occurrence of Cretaceous Calycanthaceae fossils in Brazil. In the
APG IV system The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was published ...
of 2016, Calycanthaceae is placed in the Laurales order in the
magnoliids Magnoliids (or Magnoliidae or Magnolianae) are a clade of flowering plants. With more than 10,000 species, including magnolias, nutmeg, bay laurel, cinnamon, avocado, black pepper, tulip tree and many others, it is the third-largest group of a ...
clade.


References


External links


Calycanthaceae
i
L. Watson and M. J. Dallwitz (1992 onwards). The families of flowering plants.

''Flora of North America'': Calycanthaceae



"Molecular phylogeny and intra- and intercontinental biogeography of Calycanthaceae"
{{Taxonbar, from=Q867622 Magnoliid families