Cyperus Betchei
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''Cyperus betchei'' is a sedge of the family
Cyperaceae The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus ''Carex'' w ...
that is native to Australia.


Description

The
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
sedge has a tufted habit and typically grows to a height of and produces brown flowers. The
nutlet A nut is a fruit consisting of a hard or tough nutshell protecting a kernel which is usually edible. In general usage and in a culinary sense, a wide variety of dry seeds are called nuts, but in a botanical context "nut" implies that the shell ...
is noticeably beaked. The plant has a short
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
and smooth culms that have a circular to triangular cross section and reach a height of and have a diameter of . The narrow green leaves are usually shorter than the culms and have a with of . It produces compound
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s that have four to seven primary branches that have a length of about containing loosely packed spiky clusters.


Similar species

It is very similar to ''
Cyperus angustatus ''Cyperus angustatus'' is a species of sedge that is found in both Papua New Guinea and Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (con ...
'', but differs from that species by being more robust, with broader leaves with a rough keel. Also, the darker, shinier
spikelet A spikelet, in botany, describes the typical arrangement of the flowers of grasses, sedges and some other Monocots. Each spikelet has one or more florets. The spikelets are further grouped into panicles or spikes. The part of the spikelet that ...
s are distinct and
spicate This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
, and always fall off when ripe, when in development they have a small, very acuminate beak. The winged
rachilla Rachilla may refer to the following topics in botany: * Rachilla (floral axis) A spikelet, in botany, describes the typical arrangement of the flowers of grasses, sedges and some other Monocots. Each spikelet has one or more florets. The spikele ...
projects.


Taxonomy

The species was first collected in January 1883 in
Narrabri Narrabri ( ) is a locality and seat of Narrabri Shire local government area in the North West Slopes, New South Wales, Australia on the Namoi River, northwest of Sydney. It sits on the junction of the Kamilaroi Highway and the Newell Highway. At ...
, a town in the Australian state of
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
, by the German botanist
Ernst Betche Daniel Ludwig Ernst Betche (31 December 1851, in Potsdam – 28 June 1913, in Sydney) was a German-Australian horticulturist and botanist. His mother died at his birth and he was of delicate constitution all his life. His father was sufficiently ...
. Subsequently, this
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen was kept at the
National Herbarium of Victoria The National Herbarium of Victoria ( Index Herbariorum code: MEL) is one of Australia's earliest herbaria and the oldest scientific institution in Victoria. Its 1.5 million specimens of preserved plants, fungi and algae—collectively known a ...
and ignored for over half a century, until 1936, when
Georg Kükenthal Georg Kükenthal (30 March 1864 in Weißenfels – 20 October 1955 in Coburg) was a German pastor and botanist who specialized in the field of caricology. He was the brother of zoologist Willy Kükenthal (1861–1922). From 1882 to 1885 he studie ...
first described it as a
variety Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
of ''
Cyperus angustatus ''Cyperus angustatus'' is a species of sedge that is found in both Papua New Guinea and Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (con ...
'' in Engler's ''
Das Pflanzenreich Heinrich Gustav Adolf Engler (25 March 1844 – 10 October 1930) was a German botanist. He is notable for his work on alpha taxonomy, plant taxonomy and phytogeography, such as ''Die natürlichen Pflanzenfamilien'' (''The Natural Plant Families'' ...
''. Soon afterwards, in his article 1940 ''Notes on Australian Cyperaceae, III'', published in the ''Proceedings of the Royal Society of Queensland'', the botanist
Stanley Thatcher Blake Stanley Thatcher Blake (1910 – 24 February 1973) was an Australian botanist who served as president of the Royal Society of Queensland and who was associated with the Queensland Herbarium beginning in 1945 until his death. Background Prior to ...
gave his opinion that the
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
should better be regarded as a full species, and formally promoted it to such, giving a somewhat revised and expanded treatment. The
basionym In the scientific name of organisms, basionym or basyonym means the original name on which a new name is based; the author citation of the new name should include the authors of the basionym in parentheses. The term "basionym" is used in both botan ...
is thus Kükenthal's ''C. angustatus'' var. ''betchei'', which is now regarded as a
homotypic synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In nomenclature, botanical nomenclature, a synonym is a Binomial nomenclature, scientific name that applies to a taxon that (now) goes by a different ...
. Both Kükenthal and Blake classified ''C. betchei'' in the
section Section, Sectioning or Sectioned may refer to: Arts, entertainment and media * Section (music), a complete, but not independent, musical idea * Section (typography), a subdivision, especially of a chapter, in books and documents ** Section sign ...
''pinnatae'' Kük., a section Blake writes he considers a "most difficult group". He writes that one should not consider his taxonomy to be the final say on the subject, finding the species in this group a confusing lot (Blake mentions ''C. angustatus'', ''C. betchei'', ''C. carinatus'', ''C. clarus'', ''C. dactylotes'', ''C. fulvus'', ''C. gilesii'', ''C. oxycarpus'', ''C. perangustus'' and ''C. rigidellus''), but that he was merely trying to advance our understanding of these sedges. There are two known
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
: * ''Cyperus betchei'' subsp. ''betchei'' * ''Cyperus betchei'' subsp. ''commiscens'' K.L.Wilson 1991


Distribution

In
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
it is found in the
Kimberley Kimberly or Kimberley may refer to: Places and historical events Australia * Kimberley (Western Australia) ** Roman Catholic Diocese of Kimberley * Kimberley Warm Springs, Tasmania * Kimberley, Tasmania a small town * County of Kimberley, a ...
,
Pilbara The Pilbara () is a large, dry, thinly populated region in the north of Western Australia. It is known for its Aboriginal peoples; its ancient landscapes; the red earth; and its vast mineral deposits, in particular iron ore. It is also a glo ...
and Goldfields-Esperance regions. It is also found in seasonally wet areas through the
Northern Territory The Northern Territory (commonly abbreviated as NT; formally the Northern Territory of Australia) is an states and territories of Australia, Australian territory in the central and central northern regions of Australia. The Northern Territory ...
,
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
and
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
.


Ecology

It grows in sandy-loamy soils in Western Australia. It grows in drier regions, with
wet season The wet season (sometimes called the Rainy season) is the time of year when most of a region's average annual rainfall occurs. It is the time of year where the majority of a country's or region's annual precipitation occurs. Generally, the sea ...
s.


See also

*
List of Cyperus species The genus ''Cyperus'' contains the following species recognised by ''The Plant List'' in 2015. Other species have since been considered synonyms, been newly described, or seem to have been omitted from the website database at the time. See refere ...


References

{{Taxonbar, from=Q15531140 Plants described in 1940 Flora of Western Australia Flora of the Northern Territory Flora of Queensland Flora of New South Wales betchei