Cynometra Ramiflora
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A tree in the family
Fabaceae The Fabaceae or Leguminosae,International Code of Nomenc ...
, ''Cynometra ramiflora'' is found in
mangroves A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in several ...
and flooded forests from
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
in the western Pacific west to
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
in Australia, New Guinea,
Island Southeast Asia Maritime Southeast Asia comprises the countries of Brunei, Indonesia, Malaysia, the Philippines, Singapore, and East Timor. Maritime Southeast Asia is sometimes also referred to as Island Southeast Asia, Insular Southeast Asia or Oceanic Sout ...
, and
Tropical Asia Tropical Asia refers to the entirety of the areas in Asia with a tropical climate. These areas are of geographic and economic importance due to their natural resources and biodiversity, which include many species of agricultural value. There are ...
as far west as India. Its wood is used for construction and fuel, and parts of plant are ascribed medicinal use.


Etymology

The epithet of the species, ''ramiflora'' refers to the ramiflorous inflorescences, deriving from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''rami''- (pertaining to branches) and -''florus'' (flowered).


Description

The tree grows up to 10-20 m tall. The trunks dbh is up to 60 cm, it can be
buttressed A buttress is an architectural structure built against or projecting from a wall which serves to support or reinforce the wall. Buttresses are fairly common on more ancient buildings, as a means of providing support to act against the lateral (si ...
or multistemmed, and has a red blaze (longitudinal cut exposure of bark). Leaves are compound with one, rarely two, pairs of leaflets. New leaves are pink. Lateral veins form loops well inside blade margin. Inflorescence axis is up to 20 mm long, up to 20-flowered, petal are white. Fruit is an asymmetrical, roughly globose nut, roughly 45 × 39 × 34 mm, rust brown and woody, solitary seed. Flowering while in cultivation has been recorded in August and October, fruiting has been recorded in Queensland in October and in cultivation in May. In Australia, ''C. ramiflora'' can be distinguished from other ''Cynometra'' species by the glabrous rachis and petiolules of the leaves (though these are minutely hairy or glabrescent on
Christmas Island Christmas Island, officially the Territory of Christmas Island, is an Australian external territory comprising the island of the same name. It is located in the Indian Ocean, around south of Java and Sumatra and around north-west of the ...
), the globose fruit with a small beak near the apex of the dorsal side, and by the pink new leaves.


Habitat

''Cynometra ramiflora'' grows on both rocky and sandy seashores, besides tidal rivers, on the landward side of mangrove forests, and in inland forests up to 400 m elevation. It is particularly found in environments that experience flooding or high soil moisture. It is found in primary, secondary and disturbed forests. In the native limestone forest of the island of
Saipan Saipan ( ch, Sa’ipan, cal, Seipél, formerly in es, Saipán, and in ja, 彩帆島, Saipan-tō) is the largest island of the Northern Mariana Islands, a Commonwealth (U.S. insular area), commonwealth of the United States in the western Pa ...
(largest of the
Northern Mariana Islands The Northern Mariana Islands, officially the Commonwealth of the Northern Mariana Islands (CNMI; ch, Sankattan Siha Na Islas Mariånas; cal, Commonwealth Téél Falúw kka Efáng llól Marianas), is an unincorporated territory and commonw ...
), it is co-dominant in the canopy with ''
Pisonia grandis ''Pisonia grandis'', the grand devil's-claws, is a species of flowering tree in the ''Bougainvillea'' family, Nyctaginaceae. Description The tree has broad, thin leaves, smooth bark and bears clusters of green sweet-smelling flowers that matur ...
'' and, along with ''
Guamia mariannae ''Guamia'' is a monotypic genus of flowering plants in the soursop family, Annonaceae. The sole species it contains, ''Guamia mariannae'', is endemic to the Marianas Islands The Mariana Islands (; also the Marianas; in Chamorro: ''Manislan Mar ...
'', it is most common in the understorey. At
Cape York Peninsula Cape York Peninsula is a large peninsula located in Far North Queensland, Australia. It is the largest unspoiled wilderness in northern Australia.Mittermeier, R.E. et al. (2002). Wilderness: Earth’s last wild places. Mexico City: Agrupación ...
, Queensland, and Christmas Island it grows in mangrove forest and adjacent rainforest, particularly seasonally flooded areas, at elevation from sea level to 20 m. In
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand t ...
it grows in back-mangrove forests and freshwater flooded forests.


Distribution

The species is found in New Caledonia,
Caroline Islands The Caroline Islands (or the Carolines) are a widely scattered archipelago of tiny islands in the western Pacific Ocean, to the north of New Guinea. Politically, they are divided between the Federated States of Micronesia (FSM) in the centra ...
,
Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capita ...
,
Guam Guam (; ch, Guåhan ) is an organized, unincorporated territory of the United States in the Micronesia subregion of the western Pacific Ocean. It is the westernmost point and territory of the United States (reckoned from the geographic cent ...
,
Federated States of Micronesia The Federated States of Micronesia (; abbreviated FSM) is an island country in Oceania. It consists of four states from west to east, Yap, Chuuk, Pohnpei and Kosraethat are spread across the western Pacific. Together, the states comprise a ...
, Northern Mariana Islands,
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
(Australia),
Bismarck Archipelago The Bismarck Archipelago (, ) is a group of islands off the northeastern coast of New Guinea in the western Pacific Ocean and is part of the Islands Region of Papua New Guinea. Its area is about 50,000 square km. History The first inhabitants o ...
, New Guinea, Maluku,
Palau Palau,, officially the Republic of Palau and historically ''Belau'', ''Palaos'' or ''Pelew'', is an island country and microstate in the western Pacific. The nation has approximately 340 islands and connects the western chain of the Caro ...
,
Lesser Sunda Islands The Lesser Sunda Islands or nowadays known as Nusa Tenggara Islands ( id, Kepulauan Nusa Tenggara, formerly ) are an archipelago in Maritime Southeast Asia, north of Australia. Together with the Greater Sunda Islands to the west they make up t ...
,
East Timor East Timor (), also known as Timor-Leste (), officially the Democratic Republic of Timor-Leste, is an island country in Southeast Asia. It comprises the eastern half of the island of Timor, the exclave of Oecusse on the island's north-weste ...
,
Sulawesi Sulawesi (), also known as Celebes (), is an island in Indonesia. One of the four Greater Sunda Islands, and the world's eleventh-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Mindanao and the Sulu Ar ...
, Philippines,
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and eas ...
,
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's List ...
,
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
, Christmas Island, Singapore, Malaysia, Thailand, Cambodia,
Nicobar Islands The Nicobar Islands are an archipelagic island chain in the eastern Indian Ocean. They are located in Southeast Asia, northwest of Aceh on Sumatra, and separated from Thailand to the east by the Andaman Sea. Located southeast of the Indian s ...
,
Andaman Islands The Andaman Islands () are an archipelago in the northeastern Indian Ocean about southwest off the coasts of Myanmar's Ayeyarwady Region. Together with the Nicobar Islands to their south, the Andamans serve as a maritime boundary between th ...
, Bangladesh, India, Sri Lanka. It has been
naturalized Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the in ...
in the
Society Islands The Society Islands (french: Îles de la Société, officially ''Archipel de la Société;'' ty, Tōtaiete mā) are an archipelago located in the South Pacific Ocean. Politically, they are part of French Polynesia, an overseas country of the F ...
(
Tahiti Tahiti (; Tahitian ; ; previously also known as Otaheite) is the largest island of the Windward group of the Society Islands in French Polynesia. It is located in the central part of the Pacific Ocean and the nearest major landmass is Austr ...
and others) In Queensland, it is known only from the tip of Cape York. In Myanmar it is found in the
Tanintharyi Tanintharyi or Taninthayi (widely known during the British occupation and since as Tenasserim) is a small town in Taninthayi Township, Myeik District, in the Tanintharyi Region of south-western Myanmar. It is the administrative seat for the town ...
and Ayeyarwady Regions and
Rakhine State Rakhine State (; , , ; formerly known as Arakan State) is a Administrative divisions of Myanmar, state in Myanmar (Burma). Situated on the western coast, it is bordered by Chin State to the north, Magway Region, Bago Region and Ayeyarwady R ...
.


Ecology

In
Manusela National Park Manusela National Park is located on Seram island, in the Maluku archipelago of Indonesia. It is made up of coastal forest, swamp forest, lowland and montane rainforest ecosystem types. Mount Binaiya at 3,027 meters, is the highest of the park's ...
,
Seram Seram (formerly spelled Ceram; also Seran or Serang) is the largest and main island of Maluku province of Indonesia, despite Ambon Island's historical importance. It is located just north of the smaller Ambon Island and a few other adjacent is ...
, eastern Indonesia, the fruit are eaten by ''Geoffroyus geoffroyi'', the
red-cheeked parrot The red-cheeked parrot (''Geoffroyus geoffroyi'') is a species of parrot in the family Psittaculidae found in Indonesia, Papua New Guinea and the tip of northern Australia. There are 17 subspecies currently recognized. It is a stocky short-tail ...
, while the flowers or nectar are fed on by ''Philemon subcorniculatus'', the
Seram friarbird The Seram friarbird (''Philemon subcorniculatus''), also known as the grey-necked friarbird, Ceram friarbird, grey-necked honeyeater and gray-necked honeyeater, is a species of bird in the family Meliphagidae. It is endemic to Indonesia where i ...
. The plant is a food source for the
Nolidae Nolidae is a family of moths with about 1,700 described species worldwide. They are mostly small with dull coloration, the main distinguishing feature being a silk cocoon with a vertical exit slit. The group is sometimes known as tuft moths, aft ...
moth '' Carea costiplaga'' Swinhoe, 1893.


Conservation status

The tree has been rated as having least concern conservation status by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
(IUCN). This is because even though it may be locally impacted by deforestation, it has a large and widespread population with no major threats. The species is assessed as threatened in Singapore.


Vernacular names

*''cynometra'', ''wrinkle pod mangrove'' (Australia) *''kateng'', ''kepel'', ''wunut'' (Indonesia) *''katong'', ''katong laut'' (''laut''="ocean",
Malay Malay may refer to: Languages * Malay language or Bahasa Melayu, a major Austronesian language spoken in Indonesia, Malaysia, Brunei and Singapore ** History of the Malay language, the Malay language from the 4th to the 14th century ** Indonesi ...
) *''châmpré:nh'' ( Khmer) *''myinga'', ''ye-minga'' (Myanmar)]


Uses

''Cynometra ramiflora'' has hard and heavy timber, which is used in small volumes in construction, including for door posts. The wood is also used for tool handles, wood craft and ornamental work. The species is cultivated for ornamental purposes, and the leaves, roots and seeds are harvested for use in traditional medicine. The wood is used for temporary constructions in Cambodia. It makes excellent firewood. In India, in order to treat skin diseases, the leaf is boiled in cow's milk, mixed with honey and then applied externally. Oil from the seed is also used for skin diseases, applied externally, while the root is given as a purgative and cathartic. Amongst the
Rejang people Rejang people ( Rejang: ''Tun Hejang'' or ''Tun Jang'') are an Austronesian ethnolinguistic group, native to the some parts of Bengkulu Province and South Sumatera Province in the southwestern part of Sumatera Island, Indonesia. They occupied ...
of southwestern Sumatera, ''C. ramiflora'' is one of the trees that are most frequently regarded as a ''sialang'' tree. These are trees, tall and outstanding in the forest, with a nest of the honeybee ''
Apis dorsata ''Apis dorsata'', the giant honey bee, सिङ्गुस in Nepali, is a honey bee of South and Southeast Asia, found mainly in forested areas such as the Terai of Nepal. They are typically around long. Nests are mainly built in exposed pla ...
''. Each ''sialang'' tree is held to have a sacred occupant, usually a female deity (though sometimes called ''Sernad Belelkat'') who is the owner of the bees, nest and honey. To gather the wild honey from this tree is regarded as going into the perilous realm of the spirit world, and appropriate precautions need to be taken.


References

{{Taxonbar, from=Q3008772 ramiflora Flora of Australia Flora of tropical Asia Flora of the Pacific Plants described in 1753 Taxa named by Carl Linnaeus