Cyanolanius
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The blue vanga (''Cyanolanius madagascarinus'') is a
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
species in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Vangidae. It is in the
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Cyanolanius''. It is found in
Comoros The Comoros,, ' officially the Union of the Comoros,; ar, الاتحاد القمري ' is an independent country made up of three islands in southeastern Africa, located at the northern end of the Mozambique Channel in the Indian Ocean. It ...
,
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
, and
Mayotte Mayotte (; french: Mayotte, ; Shimaore: ''Maore'', ; Kibushi: ''Maori'', ), officially the Department of Mayotte (french: Département de Mayotte), is an overseas department and region and single territorial collectivity of France. It is loc ...
, where its natural
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s are
subtropical or tropical dry forest The tropical and subtropical dry broadleaf forest is a habitat type defined by the World Wide Fund for Nature and is located at tropical and subtropical latitudes. Though these forests occur in climates that are warm year-round, and may receive ...
and
subtropical or tropical moist lowland forest Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature. Description TSMF is generally found in large, discont ...
.


Taxonomy and systematics

In 1760, the French
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published works ...
included a description of the blue vanga in his ''Ornithologie'' based on a specimen collected on the island of
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
. He used the French name ''Le pie-griesche bleu de Madagascar'' and the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
name ''Lanius Madagascariensis coeruleus''. The two stars (**) at the start of the section indicates that Brisson based his description on the examination of a specimen. Although Brisson coined Latin names, these do not conform to the binomial system and are not recognised by the
International Commission on Zoological Nomenclature The International Commission on Zoological Nomenclature (ICZN) is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 26 commissioners from 20 countries. Orga ...
. When the
Swedish Swedish or ' may refer to: Anything from or related to Sweden, a country in Northern Europe. Or, specifically: * Swedish language, a North Germanic language spoken primarily in Sweden and Finland ** Swedish alphabet, the official alphabet used by ...
naturalist
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
updated his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' for the twelfth edition in 1766, he added 240 species that had been previously described by Brisson. One of these was the blue vanga. Linnaeus included a brief description, coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Loxia madagascarina'' and cited Brisson's work. The species is now placed in the genus ''Cyanolanius'' that was introduced by the French naturalist
Charles Lucien Bonaparte Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano (24 May 1803 – 29 July 1857), was a French naturalist and ornithologist. Lucien and his wife had twelve children, including Cardinal Lucien Bonaparte. Life and career ...
in 1854.


Subspecies

Three
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised: * ''C. m. madagascarinus'' (
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1766)
: Found in northern, eastern, and western Madagascar. Clements splits this subspecies from the Comoros subspecies and calls it the Madagascar blue vanga * ''C. m. comorensis'' ( Shelley, 1894): Also known as the Comoros blue vanga, it is found on
Mohéli Mohéli , also known as Mwali, is an autonomous island that forms part of the Comoros, Union of the Comoros. It is the smallest of the three major islands in the country. It is located in the Indian Ocean off the coast of Africa and it is the sma ...
in the
Comoro Islands The Comoro Islands or Comoros ( Shikomori ''Komori''; ar, جزر القمر , ''Juzur al-qamar''; french: Les Comores) form an archipelago of volcanic islands situated off the southeastern coast of Africa, to the east of Mozambique and northw ...
. It tends to be larger than ''madagarensis'' and is also exhibits slight differences in colour. It has occasionally been considered as distinct species * ''C. m. bensoni'' Louette & Herremans, 1982: It is found on
Grande Comore Grande Comore () is an island in Comoros off the coast of Africa. It is the largest island in the Comoros nation. Most of its population is of the Comorian ethnic group. Its population is about 316,600. The island's capital is Moroni, Comoros, ...
in the Comoro Islands. It is described from only a single immature specimen, and looks very similar to ''comorensis''. Doubts have been cast on whether it is a distinctive taxon.


Description

The blue vanga is long and weighs . Adult males are bright blue in color, with a black line through the eye which extends to base of the
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
, as well as above and below the bill. It has a paler blue
eye-ring The eye-ring of a bird is a ring of tiny feathers that surrounds the orbital ring, a ring of bare skin immediately surrounding a bird's eye. The eye-ring is often decorative, and its colour may contrast with adjoining plumage. The ring of feather ...
. The tail as well as edges and tips of the wings are black in colour. The underparts and throat are bright white in colour. Females tend to be slightly duller in colour than males, with the underparts also being
off-white Shades of white are colors that differ only slightly from pure white. Variations of white include what are commonly termed off-white colors, which may be considered part of a neutral color scheme or yellow that looks like brown. In color theo ...
in colour with a pale orange or light buffy wash. The wings are slightly tinged greenish. Juveniles have a grayish-blue head, with the rest of the upperparts being dark greenish olive. Their tail is similar in appearance to an adult female's, but has a duller green-blue pattern. The throat is dirty-white in colour, the rest of the underparts being olive-buff. Colours of bare parts vary according to subspecies. ''C. m. madagarensis'' has a vivid pale or cold blue beak, with a dark tip and cutting edges in adult males, and a dark brown with similar pattern in adult female. Juveniles have a black bill with pale pink base. The iris is sky-blue or greenish blue in males, and brown in females and juveniles. The legs are black in male and gray in females. ''C. m. comorensis'' has a ultramarine blue beak with a black tip in males and a dark brown tip in females. The iris is pale blue in adults, while the legs and feet are black or dark slate in colour.


Vocalisations

Calls are described as a very characteristic ''tcccch'', which is sometimes repeated to form a ''chrr-crrk-crrk-crrrrk-crk-crk'', similar to the chabert vanga's call. The song can be faster or slower, and the first note is usually of a higher pitch. They also make a ''teea teea teea'' or a harsh ''scheet'', which is thought to be an
alarm call In animal communication, an alarm signal is an antipredator adaptation in the form of signals emitted by social animals in response to danger. Many primates and birds have elaborate alarm calls for warning conspecifics of approaching predators ...
. Race ''comorensis'' has similar calls, but also makes a grating ''erch-chhh-crkk-chh-chhh-chhh-crrk'' or a harsher ''crew-crew-crew''.''


Behaviour and ecology


Diet

Mainly feeds on
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s, including
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s,
crickets Crickets are orthopteran insects which are related to bush crickets, and, more distantly, to grasshoppers. In older literature, such as Imms,Imms AD, rev. Richards OW & Davies RG (1970) ''A General Textbook of Entomology'' 9th Ed. Methuen 8 ...
,
cicada The cicadas () are a superfamily, the Cicadoidea, of insects in the order Hemiptera (true bugs). They are in the suborder Auchenorrhyncha, along with smaller jumping bugs such as leafhoppers and froghoppers. The superfamily is divided into two ...
s,
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s, as well as
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s. They also occasionally feed upon
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
. Birds on Mohéli have been recorded eating fruit. It forages in the middle and upper levels of trees. Feeding often occurs while hanging upside down from its feet. Most food is
gleaned Gleaning is the act of collecting leftover crops from farmers' fields after they have been commercially harvested or on fields where it is not economically profitable to harvest. It is a practice described in the Hebrew Bible that became a legall ...
from the substrate, and it also sally-gleans from branches and leaves. It usually forages in groups of up to 6 birds n the non-breeding season, and in pairs while nesting. It has also been observed foraging in mixed-species flocks throughout its range.


Breeding

The breeding season is thought to be from October–December. Juveniles are taken care of by both sexes. The observed nests were placed around above the ground, near a fork between near-vertical, slender branches. The nest is cup-shaped, and made out of fine twigs and a pale down-like material. Eggs are pale blue to green and have mid-sized lilac-gray to red-brown speckles, which are the densest at the larger end. They are 19.9–21.2 × 15.8–16.2 mm in size.


Distribution

The species is found in northern, eastern, and western Madagascar, as well as on the islands of Grand Comore and Mohéli in the Comoros Islands. It inhabits the midstory and
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an a ...
of
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
dry forests in the western part of its range, and evergreen humid forest in the west. It has also been observed in littoral forests, the adjacent second growth, and even in
mangrove A mangrove is a shrub or tree that grows in coastal saline water, saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evoluti ...
s. It occurs up to an elevation of , although it has been reported up to elevations of . It inhabits a variety of woodland on the Comoro Islands. It is known to inhabits habitats above an elevation of on Mohéli. On Grand Comore, it is mainly known from degraded forest patches with banana cultivation on the slopes of
Mount Karthala Mount Karthala or Karthola ( ar, القرطالة ''Al Qirṭālah'') is an active volcano and the highest point of the Comoros at above sea level. It is the southernmost and larger of the two shield volcanoes forming Grande Comore island, the ...
at an elevation of .


Status

The nominate race on Madagascar faces few threats due to its large range and population. It is classified by the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
as least concern. However, the subspecies on the Comoro Islands is threatened by a limited range, deforestation and habitat loss, and degradation of the habitat due to invasive plant species like '' Syzygium jambos'', ''
Lantana camara ''Lantana camara'' (common lantana) is a species of flowering plant within the verbena family (Verbenaceae), native to the American tropics. It is a very adaptable species, which can inhabit a wide variety of ecosystems; once it has been introduc ...
'' and ''
Clidemia hirta ''Miconia crenata'', (syn. ''Clidemia hirta''), commonly called soapbush, clidemia or Koster's curse, is a perennial shrub. It is an invasive plant species in many tropical regions of the world, creating serious damage. Description ''Miconia cre ...
''.


References

{{Taxonbar, from=Q509183
blue vanga The blue vanga (''Cyanolanius madagascarinus'') is a bird species in the family Vangidae. It is in the monotypic genus ''Cyanolanius''. It is found in Comoros, Madagascar, and Mayotte, where its natural habitats are subtropical or tropical dry ...
Birds of the Comoros Birds of Madagascar Birds of Mayotte
blue vanga The blue vanga (''Cyanolanius madagascarinus'') is a bird species in the family Vangidae. It is in the monotypic genus ''Cyanolanius''. It is found in Comoros, Madagascar, and Mayotte, where its natural habitats are subtropical or tropical dry ...
Taxonomy articles created by Polbot Taxa named by Carl Linnaeus