Cupes
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''Cupes'' is a genus of beetles in the family
Cupedidae The Cupedidae are a small family of beetles, notable for the square pattern of "windows" on their elytra (hard forewings), which give the family their common name of reticulated beetles. The family consists of about 30 species in 9 genera, with ...
. The Cupedidae are typical “reticulate” or “net-winged” beetles with incompletely sclerotized elytra that produce the characteristic reticulate appearance. ''Cupes'' contains a single
living Living or The Living may refer to: Common meanings *Life, a condition that distinguishes organisms from inorganic objects and dead organisms ** Living species, one that is not extinct *Personal life, the course of an individual human's life * Hu ...
species '' Cupes capitatus'' and a number of
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
species described from fossils dating from the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
. ''C. capitatus'' is native to eastern North America, while the fossils are described from China and Europe. ''C. capitatus'' adults are long on average with red to gray-brown body color and a bright orange head that is molded into several knobby protuberances.Hörnschemeyer, T. 2010
''Cupes capitatus'' Fabricius 1801.
Version 17 December 2010. The Tree of Life Web Project.


Species

*'' Cupes capitatus'' Fabricius, 1801 (extant eastern North America) *†''
Cupes distinctissimus ''Cupes'' is a genus of beetles in the family Cupedidae. The Cupedidae are typical “reticulate” or “net-winged” beetles with incompletely sclerotized elytra that produce the characteristic reticulate appearance. ''Cupes'' contains a si ...
'' Kirejtshuk ''et al.'', 2016 (
Thanetian The Thanetian is, in the International Commission on Stratigraphy, ICS Geologic timescale, the latest age (geology), age or uppermost stage (stratigraphy), stratigraphic stage of the Paleocene epoch (geology), Epoch or series (stratigraphy), Serie ...
,
Menat Formation The Menat Formation is a geologic formation in France. It preserves fossils dating back to the Paleocene. It is a maar deposit located on top of an ancient volcano, the extent is very localised with the outcropping area being around 600 by 1000 ...
France) *†''
Cupes eckfeldensis ''Cupes'' is a genus of beetles in the family Cupedidae. The Cupedidae are typical “reticulate” or “net-winged” beetles with incompletely sclerotized elytra that produce the characteristic reticulate appearance. ''Cupes'' contains a si ...
'' (Tröster, 1993) (
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
,
Eckfelder Maar The Eckfelder Maar is a former volcanic lake, that was formed during the Middle Eocene around 44.3 million years ago and is thus the oldest known maar. It lies in the southwestern Eifel mountains near Manderscheid in Germany.
, Germany) *†''
Cupes golovatchi ''Cupes'' is a genus of beetles in the family Cupedidae. The Cupedidae are typical “reticulate” or “net-winged” beetles with incompletely sclerotized elytra that produce the characteristic reticulate appearance. ''Cupes'' contains a si ...
'' Kirejtshuk 2020 (Eocene, Eckfelder Maar, Germany) *†''
Cupes groehni ''Cupes'' is a genus of beetles in the family Cupedidae. The Cupedidae are typical “reticulate” or “net-winged” beetles with incompletely sclerotized elytra that produce the characteristic reticulate appearance. ''Cupes'' contains a si ...
'' Kirejtshuk, 2005 (
Priabonian The Priabonian is, in the ICS's geologic timescale, the latest age or the upper stage of the Eocene Epoch or Series. It spans the time between . The Priabonian is preceded by the Bartonian and is followed by the Rupelian, the lowest stage of t ...
,
Baltic amber The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that these forests created more than 1 ...
) *†''
Cupes hoffeinsorum ''Cupes'' is a genus of beetles in the family Cupedidae. The Cupedidae are typical “reticulate” or “net-winged” beetles with incompletely sclerotized elytra that produce the characteristic reticulate appearance. ''Cupes'' contains a si ...
'' Kirejtshuk, 2005 (Priabonian, Baltic amber) *†'' Cupes kerneggeri'' Kirejtshuk, 2005 (Priabonian, Baltic amber) *†''
Cupes komissari ''Cupes'' is a genus of beetles in the family Cupedidae. The Cupedidae are typical “reticulate” or “net-winged” beetles with incompletely sclerotized elytra that produce the characteristic reticulate appearance. ''Cupes'' contains a si ...
'' Kirejtshuk, 2005 (Priabonian, Baltic amber) *†''
Cupes legalovi ''Cupes'' is a genus of beetles in the family Cupedidae. The Cupedidae are typical “reticulate” or “net-winged” beetles with incompletely sclerotized elytra that produce the characteristic reticulate appearance. ''Cupes'' contains a si ...
'' Kirejtshuk 2020 (Eocene, Eckfelder Maar, Germany) *†''
Cupes longus ''Cupes'' is a genus of beetles in the family Cupedidae. The Cupedidae are typical “reticulate” or “net-winged” beetles with incompletely sclerotized elytra that produce the characteristic reticulate appearance. ''Cupes'' contains a si ...
'' Hong & Wang, 1987 (
Burdigalian The Burdigalian is, in the geologic timescale, an age (geology), age or stage (stratigraphy), stage in the early Miocene. It spans the time between 20.43 ± 0.05 annum, Ma and 15.97 ± 0.05 Ma (million years ago). Preceded by the Aquitanian (sta ...
, Shangwan Formation, China) *†''
Cupes lutzi ''Cupes'' is a genus of beetles in the family Cupedidae. The Cupedidae are typical “reticulate” or “net-winged” beetles with incompletely sclerotized elytra that produce the characteristic reticulate appearance. ''Cupes'' contains a si ...
'' Kirejtshuk 2020 (Eocene, Eckfelder Maar, Germany) *†'' Cupes manifestus'' Kirejtshuk ''et al.'', 2010 (Thanetian, Menat Formation, France) *†'' Cupes messelensis'' (Tröster, 1993) (
Lutetian The Lutetian is, in the geologic timescale, a stage or age in the Eocene. It spans the time between . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it is sometimes referred to as the Midd ...
,
Messel pit The Messel pit (german: Grube Messel) is a disused quarry near the village of Messel ( Landkreis Darmstadt-Dieburg, Hesse) about southeast of Frankfurt am Main, Germany. Bituminous shale was mined there. Because of its abundance of well-preserv ...
, Germany) *†'' Cupes nabozhenkoi'' Kirejtshuk 2020 (Lutetian, Messel pit, Germany) *†''
Cupes ponomarenkoi ''Cupes'' is a genus of beetles in the family Cupedidae. The Cupedidae are typical “reticulate” or “net-winged” beetles with incompletely sclerotized elytra that produce the characteristic reticulate appearance. ''Cupes'' contains a si ...
'' Kirejtshuk ''et al.'', 2010 (Ypresian,
Oise amber Oise ( ; ; pcd, Oése) is a department in the north of France. It is named after the river Oise. Inhabitants of the department are called ''Oisiens'' () or ''Isariens'', after the Latin name for the river, Isara. It had a population of 829,419 ...
, France) *†'' Cupes praeglacialis'' Gersdorf, 1976 (
Piacenzian The Piacenzian is in the international geologic time scale the upper stage (stratigraphy), stage or latest age (geology), age of the Pliocene. It spans the time between 3.6 ± 0.005 year#SI prefix multipliers, Ma and 2.588 ± 0.005 Ma (million yea ...
, Germany) *†'' Cupes rohdendorfi'' Yablokov-Khnzorian, 1960 (Priabonian, Baltic amber) *†'' Cupes simillimus'' Kirejtshuk ''et al.'', 2016 (Thanetian, France) *†'' Cupes tessellatus'' Motschulsky, 1856 (Priabonian, Baltic amber) *†'' Cupes wedmannae'' Kirejtshuk 2020 (Lutetian, Messel pit, Germany) *†'' Cupes weitschati'' Kirejtshuk, 2005 (Priabonian, Baltic amber)


References

Cupedidae Monotypic Archostemata genera {{Cupedidae-stub