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The Cuban ivory-billed woodpecker ( es, carpintero real, link=no) (''Campephilus principalis bairdii'') is a subspecies of the
ivory-billed woodpecker The ivory-billed woodpecker (''Campephilus principalis'') is a possibly extinct woodpecker that is native to the bottomland hardwood forests and temperate coniferous forests of the Southern United States and Cuba. Habitat destruction and hunting ...
native to
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...
. Originally classified as a separate species, recent research has indicated that ''C. p. bairdii'' may, in fact, be sufficiently distinct from the nominate subspecies to once again be regarded as a species in its own right. There have been no confirmed sightings of the bird since 1987; it is generally believed to be extinct, although the survival of some individuals is considered a remote possibility.


Taxonomy and appearance

''C. p. bairdii'' was originally designated as a separate species (''C. bairdii'') by
John Cassin John Cassin (September 6, 1813 – January 10, 1869) was an American ornithologist from Pennsylvania. He worked as curator and Vice President at the Philadelphia Academy of Natural Sciences and focused on the systemic classification of the Acad ...
, based on suggestions by Spencer Fullerton Baird. Cassin described it as:
Much resembling '' C. principalis'', but smaller and with the black anterior feathers of the crest longer than those succeeding, which are scarlet. White longitudinal line on the neck reaching quite to the base of the bill. ... It appears to be one of the singular insular species which have become well known to naturalists.
The Cuban form was later redesignated as a subspecies of the American ivory-billed woodpecker, ''C. principalis''. A more recent study by Fleischer, Kirchman ''et al.'' has, however, suggested that the Cuban and American forms are sufficiently genetically distinct to be regarded as separate species, which along with the
imperial woodpecker The imperial woodpecker (''Campephilus imperialis'') is a woodpecker species endemic to Mexico. If it is not extinct, it is the world's largest woodpecker species, at long. Researchers have discovered that the imperial woodpecker has slow climb ...
form a distinct North American clade within ''
Campephilus ''Campephilus'' is a genus of large American woodpeckers in the family Picidae. Taxonomy The genus ''Campephilus'' was introduced by English zoologist George Robert Gray in 1840, with the ivory-billed woodpecker (''Campephilus principalis'') as ...
'' that appeared in the Mid-
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...
. The methods adopted by the study suggest that the split between ''C. principalis'' and the lineage represented by ''C. p. bairdii'' and ''C. imperialis'' occurred first. The American Ornithologists' Union Committee on Classification and Nomenclature, while describing the data as "intriguing", has indicated that it is not yet ready to list the American and Cuban forms as two separate species. The first detailed description of the Cuban form's behavior and habitat was not published until 1893, when
Juan Gundlach Juan Cristóbal Gundlach (17 July 1810 in Marburg – 14 March 1896 in Havana), born Johannes Christoph Gundlach was a Cuban naturalist and taxonomist. Biography Gundlach graduated from Marburg University, where his father was professor of phys ...
included it in Volume 1 of his ''Ornitología Cubana''. In Cuba it was usually known by the name of ''carpintero real'' ("royal woodpecker"), although this name was also used for other birds.


Habitat

As with ''C. p. principalis'', ''C. p. bairdii'' was thought to inhabit old-growth forests with a plentiful supply of dead or dying trees; these were a source of the
cerambycid The longhorn beetles (Cerambycidae), also known as long-horned or longicorns, are a large family of beetles, with over 35,000 species described. Most species are characterized by extremely long antennae, which are often as long as or longer than ...
and other beetle larvae that formed the bulk of its diet. Most of Cuba's lowland deciduous forests had been cleared by the early 20th century, and the species became restricted to the montane
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accepts ...
forests in the northeastern part of the island. Its original range was given as through the Organ Mountains, in the lowland forests of the Ensenada de Cochinos and along the
Hanabana River Hanabana River is a river of southern Cuba. See also *List of rivers of Cuba This is an incomplete list of rivers of Cuba, arranged from west to east, by coast, with respective tributaries indented under each larger stream's name. North Coast ...
.


Behavior

Relatively few accounts of the bird's behavior in the wild exist. The ornithologist
John Dennis John Dennis may refer to: *John Dennis (dramatist) (1658–1734), English dramatist * John Dennis (1771–1806), Maryland congressman *John Dennis (1807–1859), his son, Maryland congressman *John Stoughton Dennis (1820–1885), Canadian surveyor ...
located a few birds in 1948 and noted some of their feeding and other habits, commenting that "they spent so much time reening and scratchingthat I considered it unusual". He observed that they were not especially shy or elusive once they had become used to his presence, eventually "seeming positively lethargic", although a male bird intervened quickly to drive a sparrow-hawk away from the nesting site. The breeding season of ''C. p. bairdii'' occurred from March–June. It has been surmised that the woodpecker's foraging specialization may have led to its forming small groups, much like ''Campephilus imperialis'' which was occasionally reported in groups of eight or more individuals; this behavior would enable the birds to best exploit a patchily-occurring food source.


Status

Although once common on the island, ''C. p. bairdii'' was already very rare by the late 1940s, when Dennis located a small population in a remnant of forest in the Cuchillas de Moa range which had already been cut-over for timber some years previously. George Lamb found six territories still there in 1956, and recommended that a conservation plan be implemented, but the 1959
Cuban Revolution The Cuban Revolution ( es, Revolución Cubana) was carried out after the 1952 Cuban coup d'état which placed Fulgencio Batista as head of state and the failed mass strike in opposition that followed. After failing to contest Batista in co ...
was to intervene. The last universally accepted sighting of a Cuban ivory-billed woodpecker occurred in 1987, when a single female specimen was identified in the mountains of eastern Cuba by Giraldo Alayón and Aimé Pasada, following a handful of observations of both male and female birds by a team of ornithologists, including Lester L. Short and his wife Jennifer F. M. Horne, in the area of Ojito de Agua, a hilly pine forest.Lammertinck, M
No More Hope for the Ivory-Billed Woodpecker
, Cotinga 3, 1993
Although the area was immediately designated as protected by the Cuban government, searches in 1991 and 1993 failed to find any further traces of the bird, and it became clear that the birds seen in 1986–87 had already been in "dire" circumstances. Thus the Cuban ivory-bill was inferred to have gone extinct around 1990. The area given protection in the 1980s is now part of
Alejandro de Humboldt National Park Alejandro de Humboldt National Park ( es, Parque Nacional Alejandro de Humboldt) is a national park in the Cuban provinces of Holguín and Guantánamo. It is named after the German scientist Alexander von Humboldt who visited the island in 1800 ...
. The IUCN Red List notes that calls were reportedly heard in 1998 in the highest reaches of the
Sierra Maestra The Sierra Maestra is a mountain range that runs westward across the south of the old Oriente Province in southeast Cuba, rising abruptly from the coast. The range falls mainly within the Santiago de Cuba and in Granma Provinces. Some view it a ...
, but that a subsequent search failed to find any trace of the species or of good potential habitat: it is considered a (remote) possibility that some individuals may survive, as around 80% of suitable habitat in Cuba has yet to be searched.The Sierra Maestra was not part of the species' recorded range.


References

{{Taxonbar, from=Q1121554 Cuban ivory-billed woodpecker Endemic birds of Cuba Cuban ivory-billed woodpecker Cuban ivory-billed woodpecker Articles containing video clips