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The Cuban green woodpecker (''Xiphidiopicus percussus'') is a species of
woodpecker Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions. ...
in the family
Picidae Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions. M ...
and tribe Melanerpini, known locally in
Cuban Spanish Cuban Spanish is the variety of the Spanish language as it is spoken in Cuba. As a Caribbean variety of Spanish, Cuban Spanish shares a number of features with nearby varieties, including coda weakening and neutralization, non-inversion o ...
as carpintero verde (literally "green woodpecker"). It is the only species within the genus ''Xiphidiopicus'' and is one of two woodpeckers
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...
. It is the most widespread and common woodpecker in Cuba, inhabiting primarily woodlands, as well as dry and wet forests, pine forests and
mangrove A mangrove is a shrub or tree that grows in coastal saline water, saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evoluti ...
s. The population of the Cuban green woodpecker is stable and its status is listed as "Least Concern".


Appearance

A medium-sized and colorful bird, the Cuban green woodpecker is similar to a
sapsucker The sapsuckers are species of North American woodpeckers in the genus ''Sphyrapicus''. Taxonomy and systematics The genus ''Sphyrapicus'' was introduced in 1858 by the American naturalist Spencer Baird with the yellow-bellied sapsucker (''Sphyra ...
in general shape and size. Featuring olive green overparts and yellow, streaked underparts, the Cuban green woodpecker usually appears crested, and features a bright red upper breast, as well as a black chin contrasting a white face and
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also ...
. In males, a red
crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...
is present, while females possess black crowns with white stripes. The area around the
cloaca In animal anatomy, a cloaca ( ), plural cloacae ( or ), is the posterior orifice that serves as the only opening for the digestive, reproductive, and urinary tracts (if present) of many vertebrate animals. All amphibians, reptiles and birds, a ...
, or the crissum, is yellow with black barring. Its beak is short and straight with a progressively darkening coloration, from dark blue to black from base to tip dark blue to black coloration. They have been shown to have brown irises and grayish green tarsi and toes Cuban green woodpeckers show slight sexually dimorphic qualities; females have shorter bills and are generally significantly smaller than their male counterparts. Juveniles will show more barring and streaking on their breasts and lack the depth of color adults have in their
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
. They typically measure between in length and weigh approximately .


Taxonomy

The Cuban green woodpecker as a species is unique in both genetics and appearance. Historically, it was presumed that the Cuban green woodpecker (''Xiphidiopicus percussus)'', and the
Hispaniolan woodpecker The Hispaniolan woodpecker (''Melanerpes striatus'') is a medium-sized woodpecker endemic to the Caribbean island of Hispaniola (split between the Dominican Republic and Haiti). Description The Hispaniolan woodpecker is a gold and black barred ...
(''Melanerpes striatus'') were sister taxa and there was speculation that it had relationships to the genera ''
Sphyrapicus The sapsuckers are species of North American woodpeckers in the genus ''Sphyrapicus''. Taxonomy and systematics The genus ''Sphyrapicus'' was introduced in 1858 by the American naturalist Spencer Baird with the yellow-bellied sapsucker (''Sphyr ...
'' and ''
Melanerpes ''Melanerpes'' is a genus of woodpeckers of the family Picidae found in the New World. The 24 members of the genus are mostly colourful birds, conspicuously barred in black and white, with some red and yellow. Taxonomy The genus ''Melanerpes'' ...
''. According to research done in 2006, "''X. percussus'' is the sister taxon to the ''Melanerpes'' woodpeckers, which appear to group into a single distinct clade." It was also found that ''Xiphidiopicus percussus'' is not the sister taxon to ''M. striatus'' and that the genus ''Sphyrapicus'' diverged earlier than ''Xiphidiopicus,'' which diverged during the late
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
-early
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
or
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. In terms of appearance, no other Cuban species of woodpecker unmarked green upperparts. The
Yellow-bellied Sapsucker The yellow-bellied sapsucker (''Sphyrapicus varius'') is a medium-sized woodpecker that breeds in Canada and the northeastern United States. Taxonomy The yellow-bellied sapsucker was described and illustrated using a hand-coloured plate by the E ...
(''Sphyrapicus varius'') is similar in coloration and also has a facial pattern that is reminiscent to that of Cuban green woodpecker; however, the sapsucker's upperparts are white and black and has a broad black band across its breast.


Subspecies

The species can be divided into the following subspecies, with only two being formerly recognized: * ''X. p. insulaepinorum -'' found in the Cantiles Keys, Isla de la Juventud and Archipiélago Jardines de la Reina (Cayo Caballones). * ''X. p. percussus'' - found on the archipelagos of Sabana and Camagüey * ''X. p. cocoensis'' * ''X. p. gloriae'' * ''X. p. marthae'' * ''X. p. monticola''


Habitat and distribution

The Cuban green woodpecker is only found in Cuba, but is extremely common on the island. Its natural
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s are wet and
dry forests The tropical and subtropical dry broadleaf forest is a habitat type defined by the World Wide Fund for Nature and is located at tropical and subtropical latitudes. Though these forests occur in climates that are warm year-round, and may receive ...
, mangroves, open woodland with palms, and pine forests, lowland moist forests, and heavily degraded former forest.


Behavior

The Cuban green woodpecker is found most frequently in pairs, and less frequently in small groups. It is able to remain completely still for long periods and will forage for dead insects on dead limbs at any height within its habitat. They are very territorial, especially during breeding season, and use active vocalization, utilizing both drumming and vocal signals, and active pursuit, pursuing threats while flicking their wings, in order deter intruders in their nests, which is usually the
West Indian woodpecker The West Indian woodpecker (''Melanerpes superciliaris'') is a species of bird in subfamily Picinae of the woodpecker family Picidae. It is found in the Bahamas, the Cayman Islands and Cuba. Taxonomy and systematics The West Indian woodpecke ...
(''Melanerpes superciliaris''). Similarly to all other woodpeckers, its flight is undulating.


Vocalizations


Calls and songs

The Cuban green woodpecker's call is a short, harsh "''jhhhorr, jhhhorr, jhhhorr,''" as well as a higher pitched "''yeh-yeh-yeh''." The call is usually single noted with double notes occasionally being uttered. It may sometimes produce a slurred, two-note call “ta-ha”, which is similar to the
Yellow-bellied Sapsucker The yellow-bellied sapsucker (''Sphyrapicus varius'') is a medium-sized woodpecker that breeds in Canada and the northeastern United States. Taxonomy The yellow-bellied sapsucker was described and illustrated using a hand-coloured plate by the E ...
(Sphyrapicus varius)'s call. It will use its call mostly while protecting nest and chicks from intruder.


Diet

The Cuban green woodpecker's is primarily an
insectivore A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores wer ...
, focusing on large insects, but will also consume
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
s and small
frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...
s occasionally. It has also been documented taking eggs of other birds such as
heron The herons are long-legged, long-necked, freshwater and coastal birds in the family Ardeidae, with 72 recognised species, some of which are referred to as egrets or bitterns rather than herons. Members of the genera ''Botaurus'' and ''Ixobrychus ...
s nesting in mangroves. It will also feed on flower
nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...
, especially ''
Cordia sebestena ''Cordia sebestena'' is a shrubby tree in the borage family, Boraginaceae, native to the Americas, American tropics. It ranges from southern Florida in the United States and the Bahamas, southwards throughout Central America and the Greater Anti ...
''.Dalsgaard, B. (2011). Nectar-feeding and pollination by the Cuban Green Woodpecker (Xiphidiopicus percussus) in the West Indies. ''Ornitología Neotropical'', ''22'', 447-451. While perching itself on the plant or on a nearby branch to reach the flowers, it will insert its bill into the
floral tube In angiosperms, a hypanthium or floral cup is a structure where basal portions of the Sepal, calyx, the petal, corolla, and the stamens form a cup-shaped tube. It is sometimes called a floral tube, a term that is also used for corolla tube and cal ...
to drink the nectar for a few seconds, by doing so its bill will come into contact with
anthers The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
and stigma inside the floral tube, making the bird then act as a
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the maj ...
. Males and females utilize different ways to exploit food sources, as the male is larger than the female.


Reproduction


Courtship

There is very little known about the courtship of the Cuban green woodpecker. Usually, woodpeckers perform flutter-aerial-display including a
gliding flight Gliding flight is heavier-than-air flight without the use of thrust; the term volplaning also refers to this mode of flight in animals. It is employed by gliding animals and by aircraft such as gliders. This mode of flight involves flying a s ...
with wings held well above the back, accompanied by calls.
Courtship Courtship is the period wherein some couples get to know each other prior to a possible marriage. Courtship traditionally may begin after a betrothal and may conclude with the celebration of marriage. A courtship may be an informal and private m ...
feeding by male to female most likely occurs as well. They are probably monogamous.


Breeding season, nesting and hatching

Cuban green woodpeckers will nest in the cavities of tree, alive or dead, often 4–5 metres above the ground. The male will usually do most of the excavation for the nest-hole, and will endure the nesting duties with a relatively high contribution. The breeding season takes place between February and August, with peak in May–June, probably associated with rains.Kirkconnell, A. (2000). Variación morfológica del Carpintero Verde Xiphidiopicus percussus en Cuba. Cotinga. 14: 94–98.http://www.neotropicalbirdclub.org/wp-content/uploads/2016/05/Cotinga-14-2000-94-98.pdf The female will lay 3-4 white eggs and both adults share the incubation, which is presumed to be between 9 and 14 days, however more information is currently lacking, and feed the chicks, with the females feeding the nestlings at a higher rate. The chicks are hatched completely naked and stay in the nest for about 3–4 weeks. They are sexually mature in the first year of life.


References


External links

* * * * * * * {{Taxonbar, from=Q886565 Cuban green woodpecker Endemic birds of Cuba Higher-level bird taxa restricted to the West Indies Cuban green woodpecker Taxonomy articles created by Polbot Taxa named by Coenraad Jacob Temminck