The Cuban cactus scrub is a

xeric shrubland Deserts and xeric shrublands are a biome defined by the World Wide Fund for Nature. Deserts and xeric (ancient Greek xērós, “dry") shrublands form the largest terrestrial biome, covering 19% of Earth's land surface area. Ecoregions in this h ...

ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of l ...

that occupies on the

leeward Windward () and leeward () are terms used to describe the direction of the wind. Windward is ''upwind'' from the point of reference, i.e. towards the direction from which the wind is coming; leeward is ''downwind'' from the point of reference ...

coast of

Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...

. Most of it occurs in the southeastern part of the island in the provinces of

Guantánamo Guantánamo (, , ) is a municipality and city in southeast Cuba and capital of Guantánamo Province. Guantánamo is served by the Caimanera port near the Guantanamo Bay Naval Base, site of a U.S. naval base. The area produces sugarcane and cotton ...

and

Santiago de Cuba Santiago de Cuba is the second-largest city in Cuba and the capital city of Santiago de Cuba Province. It lies in the southeastern area of the island, some southeast of the Cuban capital of Havana. The municipality extends over , and contains t ...

. The ecoregion receives less than of rainfall annually. The principal

soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former te ...

s are coastal

rendzina Rendzina (or ''rendsina'') is a soil type recognized in various soil classification systems, including those of Britain and Germany as well as some obsolete systems. They are humus-rich shallow soils that are usually formed from carbonate- or occ ...

s that were derived from

coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and sec ...

line

limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...

. Cuban cactus scrub contains four vegetation zones:

xerophytic A xerophyte (from Greek ξηρός ''xeros'' 'dry' + φυτόν ''phuton'' 'plant') is a species of plant that has adaptations to survive in an environment with little liquid water, such as a desert such as the Sahara or places in the Alps or th ...

coastal and subcoastal scrubland, coastal thorny semidesert, coastal

sclerophyll Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short internodes (the distance between leaves along the stem) and leaf orientation which is parallel or oblique to direct ...

ous scrubland, and rocky coastal scrublands.

Xerophytic coastal and subcoastal scrubland

Vegetation in the xerophytic coastal and subcoastal scrublands reaches a height of and is dominated by palms and

succulent In botany, succulent plants, also known as succulents, are plants with parts that are thickened, fleshy, and engorged, usually to retain water in arid climates or soil conditions. The word ''succulent'' comes from the Latin word ''sucus'', meani ...

s, especially

cacti A cactus (, or less commonly, cactus) is a member of the plant family Cactaceae, a family comprising about 127 genera with some 1750 known species of the order Caryophyllales. The word ''cactus'' derives, through Latin, from the Ancient Greek ...

. Common evergreen plants include (''

Bourreria ''Bourreria'' is a genus of flowering plants in the borage family, Boraginaceae. Members of the genus are commonly known as strongbark or strongback. The generic name was chosen by Patrick Browne to honour German pharmacist Johann Ambrosius Beu ...

virgata''), ('' Capparis cynophallophora''), ('' Eugenia foetida''), ''

Bursera ''Bursera'' is a genus with about 100 described species of flowering shrubs and trees varying in size up to high. It is the type genus for Burseraceae. The trees are native (often for many species endemic) to the Americas, from the southern Uni ...

glauca'', '' B. cubana'', '' Croton'' spp., ''

Cordia ''Cordia'' is a genus of flowering plants in the borage family, Boraginaceae. It contains about 300 species of shrubs and trees, that are found worldwide, mostly in warmer regions. Many of the species are commonly called manjack, while ''bocote' ...

'' spp., ''

Calliandra ''Calliandra'' is a genus of flowering plants in the pea family, Fabaceae, in the mimosoid clade of the subfamily Caesalpinioideae. It contains about 140 species that are native to tropical and subtropical regions of the Americas. Description T ...

colletioides'', ''

Caesalpinia ''Caesalpinia'' is a genus of flowering plants in the family Fabaceae. Historically, membership within the genus has been highly variable, with different publications including anywhere from 70 to 165 species, depending largely on the inclusion o ...

'' spp., ''

Acacia ''Acacia'', commonly known as the wattles or acacias, is a large genus of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa and Australasia. The genus na ...

'' spp., '' Phyllostylon brasiliense'', '' Pseudosamanea cubana'' and (''

Guaiacum officinale ''Guaiacum officinale'', commonly known as roughbark lignum-vitae, guaiacwood or gaïacwood, is a species of tree in the caltrop family, Zygophyllaceae, that is native to the Caribbean and the northern coast of South America. Description This sm ...

''). (''

Opuntia stricta ''Opuntia stricta'' is a species of large cactus that is endemic to the subtropical and tropical coastal areas of the Americas, especially around the Caribbean. Common names include erect prickly pear and nopal estricto (Spanish). The first desc ...

''), '' O. militaris'', ('' Harrisia eriophora''), ('' H. taetra''), ('' Pilosocereus polygonus''), ('' Dendrocereus nudiflorus''), (''

Agave ''Agave'' (; ; ) is a genus of monocots native to the hot and arid regions of the Americas and the Caribbean, although some ''Agave'' species are also native to tropical areas of North America, such as Mexico. The genus is primarily known for ...

'' spp.), (''

Melocactus ''Melocactus'' (melon cactus), also known as the Turk's cap cactus, is a genus of cactus with about 30–40 species. They are native to the Caribbean, western Mexico through Central America to northern South America, with some species along the A ...

'' spp.) and ''

Leptocereus ''Leptocereus'' is a genus of cacti native to the Greater Antilles. It has been placed in the tribe Leptocereeae or in a broadly defined Echinocereeae The Echinocereeae are a tribe of cacti in the subfamily Cactoideae. Since 2006, the tribe ...

'' spp. are typical

Coastal thorny semidesert

Coastal thorny semidesert is similar to xerophytic coastal and subcoastal scrubland in that they both have the same succulent composition and reach a height of . However, succulents represent a smaller proportion of the plant life in this zone. Other plant species include ('' Cordia sebestena''), ('' Hippomane mancinella'') and (''

Plumeria ''Plumeria'' (), known as frangipani, is a genus of flowering plants in the subfamily Rauvolfioideae, of the family Apocynaceae. Most species are deciduous shrubs or small trees. The species variously are endemic to Mexico, Central America, and ...

filifolia'').

Coastal sclerophyllous scrubland

The vegetation in this zone is

ous, meaning that their

leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...

are hard. It reaches a height of with emergent

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

s up to . This zone represents the transition between xeric shrublands and moister

dry forests The tropical and subtropical dry broadleaf forest is a habitat type defined by the World Wide Fund for Nature and is located at tropical and subtropical latitudes. Though these forests occur in climates that are warm year-round, and may receive ...

. Plant life includes ('' Picrodendron baccatum''), (''

Maytenus ''Maytenus'' ''Sunset Western Garden Book,'' 1995:606–607 is a genus of flowering plants in the family Celastraceae. Members of the genus are distributed throughout Central and South America, Southeast Asia, Micronesia and Australasia, the Indi ...

buxifolia''), (''

Pictetia ''Pictetia'' is a genus of about eight species of trees and shrubs in the legume family with Thorns, spines, and prickles, spiny stems and (in six of the eight species) spine-tipped leaflets.Beyra & Lavin (1999), pp.36–38 The genus is endemism, ...

spinosa''), ('' Brya ebenus'') and (''

Diospyros ''Diospyros'' is a genus of over 700 species of deciduous and evergreen trees and shrubs. The majority are native to the tropics, with only a few species extending into temperate regions. Individual species valued for their hard, heavy, dark tim ...

grisebachii'').

Rocky coastal scrublands

The most sparse and stunted vegetation is found in this zone, which occurs adjacent to the ocean on

karst Karst is a topography formed from the dissolution of soluble rocks such as limestone, dolomite, and gypsum. It is characterized by underground drainage systems with sinkholes and caves. It has also been documented for more weathering-resistant ro ...

ic soil. The flora is tolerant of wind and

salt spray Salt is a mineral composed primarily of sodium chloride (NaCl), a chemical compound belonging to the larger class of salts; salt in the form of a natural crystalline mineral is known as rock salt or halite. Salt is present in vast quantiti ...

, and includes ('' Borrichia arborescens''), (''

Argusia ''Tournefortia'', commonly known as soldierbush, is a genus of flowering plants in the borage family, Boraginaceae. It was first published under the name ''Pittonia'' by Charles Plumier in 1703, in honour of Joseph Pitton de Tournefort. Later, ...

gnaphalodes''), and (''

Sesuvium ''Sesuvium'' is a genus of flowering plants in the ice plant family, Aizoaceae. The roughly eight species it contains are commonly known as sea-purslanes. Selected species * '' Sesuvium crithmoides'' Welw. – Tropical sea-purslane * '' Ses ...

maritimum'').

Fauna

Bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

s of the cactus scrub include the

Zapata sparrow The Zapata sparrow (''Torreornis inexpectata'') is a medium-sized grey and yellow bird that lives in the grasslands of the Zapata Swamp and elsewhere on the island of Cuba. Measuring about in length, it is grey and yellow overall with a dark red ...

(''Torreornis inexpectata sigmani''), Cuban vireo (''Vireo gundlachii''), Cuban gnatcatcher (''Polioptila lembeyei''), and Oriente warbler (''Teretistris fornsi''). The

yellow-striped pygmy eleuth The yellow-striped pygmy eleuth (''Eleutherodactylus limbatus''), also known as the yellow-striped dwarf frog, is a species of frog in the family Eleutherodactylidae from closed mesic and xeric forests in Cuba. The yellow-striped pygmy eleuth i ...

(''Eleutherodactylus limbatus'') is a species of

frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

to this ecoregion, while several anole species and the Cuban rock iguana ('' Cyclura nubila nubila'') are endemic

lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...

References

{{reflist Deserts and xeric shrublands Ecoregions of the Caribbean Ecoregions of Cuba * Natural history of Cuba Neotropical ecoregions