Ctenopterella Thwaitesii
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''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynesia.


Description

The rhizomes are dorsiventral (having upper and lower surfaces clearly distinct in appearance), and bear two rows of stipes, which sometimes have distinct joints where they attach. Distinct phyllopodia are present in some below the joint. The rhizome scales are brown, glabrous, and dull to glossy. Hairs, where present, are unbranched and branched, from whitish to brown in color. The leaf blades range from pinnatifid to pinnate-pinnatifid in cutting, bearing free veins which usually end in hydathodes on the upper surface of the leaf. Sori are borne beneath pinnae or lobes, sometimes slightly sunken into the leaf tissue, usually in two rows but rarely in one. The sori are circular to elliptic in shape; the sporangia lack hairs.


Taxonomy

The genus was first described by Barbara Parris in 2007 to receive some of the species of the polyphyletic genus ''
Ctenopteris ''Ctenopteris'' is a defunct genus of ferns in the family Polypodiaceae. The name is derived from two Greek words: ''ktenos'', "comb", and ''pteris'', "fern".Umberto Quattrocchi. 2000. ''CRC World Dictionary of Plant Names'' volume I. CRC Press: B ...
''; ''Ctenopterella'' is a diminutive form of that name. She initially placed twelve species in the genus. In 2013, she described a new species, ''
Ctenopterella gabonensis ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'', from the Monts Doudou in Gabon, and transferred a Vietnamese species, ''
Ctenopterella nhatrangensis ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'', into the genus. In 2015, she transferred three more species from ''Ctenopteris'', including one she had formerly placed in synonymy, in preparation for a monograph on the genus. The only phylogenetic study so far to include any ''Ctenopterella'' species sampled ''Ctenopterella denticulata'' only. It found that this species, ''
Acrosorus friderici-et-pauli ''Acrosorus friderici-et-pauli'' is a species of fern in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is native to Thailand, peninsular Malaysia, Vietnam, th ...
'' (the type of its genus), and ''
Grammitis stenophylla '' Grammitis stenophylla'', commonly known as the narrow-leafed finger fern, is a fern in the family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affin ...
'' form a clade sister to the combined clade of '' Oreogrammitis'', '' Prosaptia'', '' Radiogrammitis'', and '' Themelium''. None of the three taxa in the first clade are particularly close morphologically. ''Ctenopterella denticulata'' has since been transferred to the genus '' Chrysogrammitis'' as '' Chrysogrammitis denticulata''.


Species

, the ''Checklist of Ferns and Lycophytes of the World'' accepted the following species and hybrids: *''
Ctenopterella blechnoides ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' (Grev.) Parris *''
Ctenopterella boivinii ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' (Mett. ex Kuhn) Rakotondr. & Parris *''
Ctenopterella cornigera ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' (Baker) Parris *''
Ctenopterella fluvialis ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' Parris *''
Ctenopterella gabonensis ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' Parris *''
Ctenopterella gordonii ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' (S.B.Andrews) Parris *''
Ctenopterella khaoluangensis ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' (Tagawa & K.Iwats.) Parris *''
Ctenopterella lasiostipes ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' (Mett.) Parris *''
Ctenopterella lastii ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' Parris & Rakotondr. *''
Ctenopterella lepida ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' (Brause) Parris *''
Ctenopterella macrorhyncha ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Po ...
'' (Baker) Parris *''
Ctenopterella nhatrangensis ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' (C.Chr. & Tardieu) Parris *''
Ctenopterella pacifica ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polyne ...
'' Parris *''
Ctenopterella parvula ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polyne ...
'' (Bory ex Willd.) Parris *''
Ctenopterella pediculata ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' (Baker) Parris *''
Ctenopterella seemannii ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polyne ...
'' (J.Sm.) Parris *'' Ctenopterella thwaitesii'' (Bedd.) Parris *''
Ctenopterella undosa ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' (Baker) Parris *'' Ctenopterella vodonaivalui'' (Brownlie) Parris *''
Ctenopterella zenkeri ''Ctenopterella'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Africa through southeast Asia and Oceania to Polynes ...
'' (Hieron.) Parris


References

{{Taxonbar, from=Q17187669 Polypodiaceae Fern genera