Cteniogenys
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''Cteniogenys'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
choristodere Choristodera (from the Greek χωριστός ''chōristos'' + δέρη ''dérē'', 'separated neck') is an extinct order of semiaquatic diapsid reptiles that ranged from the Middle Jurassic, or possibly Triassic, to the late Miocene (168 to ...
, a morphologically diverse group of aquatic reptiles. It is part of the monotypic family Cteniogenidae. The type and only named species, ''C. antiquus'', was named in 1928 by
Charles W. Gilmore Charles Whitney Gilmore (March 11, 1874 – September 27, 1945) was an American paleontologist who gained renown in the early 20th century for his work on vertebrate fossils during his career at the United States National Museum (now the N ...
. The holotype, VP.001088, was collected in the
Morrison Formation The Morrison Formation is a distinctive sequence of Late Jurassic, Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandsto ...
(Como Bluff),
Wyoming Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the south ...
in 1881 by William H. Reed. More specimens have been discovered since then, including specimens from the Late Jurassic of Portugal and Middle Jurassic of Britain, which have not been assigned to species.


Description

''Cteniogenys'' was long, and probably weighed less than . The skull of this genus was long and slender, and the jaws had numerous conical teeth. ''Cteniogenys'' in the Morrison probably fed on insects and small fish. It is mostly known from freshwater sites (rivers and ponds), and is a rare find in the formation (only 60 specimens out of over 2,800 total vertebrate specimens known from the formation), mostly known from northern outcrops (particularly
Wyoming Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the south ...
); this may reflect a preservation bias against small animals in terrestrial settings, rather than an accurate reflection of ''Cteniogenys'' populations in the Morrison.


Distribution

Fossils of ''Cteniogenys'' are known from the upper Middle Jurassic (
Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Strat ...
) aged
Forest Marble Formation The Forest Marble is a geological formation in England. Part of the Great Oolite Group, it dates to the late Bathonian stage of the Middle Jurassic.Weishampel, David B; et al. (2004). "Dinosaur distribution (Middle Jurassic, Europe)." In: Weisham ...
and
Kilmaluag Formation The Kilmaluag Formation is a Middle Jurassic geologic formation in Scotland. It was formerly known as the Ostracod Limestone for the abundance of fossil freshwater ostracods within it. The Kilmaluag Formation is very fossiliferous, with ostracods ...
of Britain, the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
(
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
) aged
Alcobaça Formation The Alcobaça Formation, previously known as the Guimarota Formation and also known as the Consolação Unit, is a geological formation in Portugal. It dates back to the Kimmeridgian stage of the Late Jurassic. It is an important source of inform ...
of Portugal, and the Late Jurassic-age
Morrison Formation The Morrison Formation is a distinctive sequence of Late Jurassic, Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandsto ...
of western
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, indeterminate similar remains are also known from the
Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 166.1 ± 4.0 Ma (million years ago) and 163.5 ± 4.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
aged
Balabansai Formation The Balabansai Formation (sometimes transcribed Balabansay, also known as Balabansai Svita) is a geological formation in Kazakhstan, Kyrgyzstan and Uzbekistan whose strata date back to the Bathonian and Callovian stages of the Middle Jurassic. Di ...
of
Kyrgyzstan Kyrgyzstan,, pronounced or the Kyrgyz Republic, is a landlocked country in Central Asia. Kyrgyzstan is bordered by Kazakhstan to the north, Uzbekistan to the west, Tajikistan to the south, and the People's Republic of China to the east. ...
and the Bathonian aged
Itat Formation The Itat Formation (Russian: итатская свита) is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from t ...
of western Siberia, and the Berriasian aged
Angeac-Charente bonebed The Angeac-Charente bonebed is a fossil deposit located near Angeac-Charente in western France. It dates to the Berriasian stage of the Early Cretaceous, and is coeval with the Purbeck Group of Southern England. It has amongst the most diverse assem ...
in France.Ronan Allain, Romain Vullo, Lee Rozada, Jérémy Anquetin, Renaud Bourgeais, et al.
Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J/K boundary
Geodiversitas, Museum National d’Histoire Naturelle Paris, In press. ffhal-03264773f
A handful of skull and jaw fragments from the Late Cretaceous
Oldman Formation The Oldman Formation is a stratigraphic unit of Late Cretaceous (Campanian stage) age that underlies much of southern Alberta, Canada. It consists primarily of sandstones that were deposited in fluvial channel and floodplain environments. It was ...
and
Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...
in Canada were assigned to ''Cteniogenys'' by Gao and Fox (1998). Given the long gap in time, however, Gao et al. (2005) cautioned that these remains could constitute a different, as-yet undetermined genus. In their description of new material of ''
Khurendukhosaurus ''Khurendukhosaurus'' is a genus of choristodere, a type of amphibious reptile. It is known from Lower Cretaceous rocks of Mongolia and Russia. Two species have been named. The type species, ''K. orlovi'', was named in 1984 by Sigogneau–Rus ...
'', Matsumoto et al. (2009) agreed that the putative Cretaceous occurrence of ''Cteniogenys'' did not belong to that genus and represents an indeterminate choristoderan, possibly a new genus.


Systematics

The genus was named in 1928 by
Charles W. Gilmore Charles Whitney Gilmore (March 11, 1874 – September 27, 1945) was an American paleontologist who gained renown in the early 20th century for his work on vertebrate fossils during his career at the United States National Museum (now the N ...
on the basis of a lower jaw collected during the late 19th century by
Othniel Charles Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of Paleontology in Yale College and President of the National Academy of Sciences. He was one of the preeminent scientists in the field of paleontology. Among h ...
's workers at
Como Bluff Como Bluff is a long ridge extending east–west, located between the towns of Rock River and Medicine Bow, Wyoming. The ridge is an anticline, formed as a result of compressional geological folding. Three geological formations, the Sundance, t ...
. He tentatively described the genus as a lizard, noting that it could instead be a frog. New material of ''Cteniogenys'' from Europe prompted
Susan E. Evans Susan E. Evans is British palaeontologist and herpetologist. She is the author or co-author of over 100 peer-reviewed papers and book chapters. She received a BSc in Zoology at Bedford College in 1974, and in 1977 a PhD in vertebrate palaeon ...
to reclassify it as an early choristodere in 1989. To date it is only known from fragmentary specimens. In phylogenic analyses it has been found to the basal most choristodere.


See also

*
Paleobiota of the Morrison Formation The Morrison Formation is a distinctive sequence of Late Jurassic sedimentary rock that is found in the western United States, which has a wide assortment of Taxon, taxa represented in its fossil record, including dinosaur fossils in North America ...


References

Choristodera Jurassic reptiles Late Jurassic reptiles of North America Jurassic Colorado Jurassic geology of Oklahoma Jurassic geology of South Dakota Jurassic geology of Utah Jurassic geology of Wyoming Late Cretaceous reptiles of North America Fossil taxa described in 1928 Prehistoric reptile genera {{cretaceous-reptile-stub