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''Cryptocarya'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s belonging to the laurel family,
Lauraceae Lauraceae, or the laurels, is a plant family that includes the true laurel and its closest relatives. This family comprises about 2850 known species in about 45 genera worldwide (Christenhusz & Byng 2016 ). They are dicotyledons, and occur ma ...
. The genus includes more than 350 species, distributed through the
Neotropical The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
,
Afrotropical The Afrotropical realm is one of Earth's eight biogeographic realms. It includes Africa south of the Sahara Desert, the majority of the Arabian Peninsula, the island of Madagascar, southern Iran and extreme southwestern Pakistan, and the island ...
,
Indomalayan The Indomalayan realm is one of the eight biogeographic realms. It extends across most of South and Southeast Asia and into the southern parts of East Asia. Also called the Oriental realm by biogeographers, Indomalaya spreads all over the Indi ...
, and
Australasian realm The Australasian realm is a biogeographic realm that is coincident with, but not (by some definitions) the same as, the geographical region of Australasia. The realm includes Australia, the island of New Guinea (comprising Papua New Guinea and ...
s.


Overview

The genus includes
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of evergreen trees, distributed mostly in tropical and subtropical regions of South America, India, China, Java, New Guinea, Africa, Madagascar, and Mauritius, with seven species in Southern Africa. Common in the canopy, they grow up to 60 m, or as subcanopy trees in the succession climax species in tropical, lower temperate, or subtropical broadleaved forests. They are found in low-elevation evergreen forests and littoral rainforests, on all type of soils. The seeds are readily dispersed by fruit-eating birds, and seedlings and saplings have been recorded from other habitats where they are unlikely to develop to maturity. The genus name ''Cryptocarya'' is from a Greek word ''krypto'' meaning to hide, ''karya'' meaning a walnut tree, the fruit of which was known as ''karyon'', a word also used to describe other fruits. Sometimes, they are called mountain laurels or mountain walnuts. The fruit are succulent, partially immersed in a deep, thick cup. The species forming this genus share a unique paniculate inflorescence with the ultimate divisions that are not quite cymose; that is, the lateral flowers of what looks like a cyme are not strictly opposite, but tend to be subopposite, while in most genera of Lauraceae with paniculate inflorescences the lateral flowers in a cyme are strictly opposite. In a recent generic classification of Lauraceae based on DNA sequence, ''Cryptocarya'' was found to be part of a strongly supported clade that also includes ''
Beilschmiedia ''Beilschmiedia'' is a genus of trees and shrubs in family Lauraceae. Most of its species grow in tropical climates, but a few of them are native to temperate regions, and they are widespread in tropical Asia, Africa, Madagascar, Australia, N ...
, Potameia,
Endiandra ''Endiandra'' is a genus of about 126 species of plants, mainly trees, in the laurel family Lauraceae. They are commonly called "walnut" despite not being related to the Northern Hemisphere walnuts (''Juglans'' spp.) which are in the family Jugl ...
'' and ''
Aspidostemon ''Aspidostemon'' is a genus of flowering plants belonging to the family Lauraceae. It occurs in Madagascar. Taxonomy The genus was described by Jens Gunter Rohwer & Hans Georg Richter in ''Jahrbuch für Botanische Systematik, Pflanzengeschich ...
''. In both ''Aspidostemon '' and ''Cryptocarya'', the fruit are enclosed in the enlarged hypanthium, but this might be a parallel development and not a signal of common ancestry.


Ecology

The ecological requirements of the genus are those of the
laurel forest Laurel forest, also called laurisilva or laurissilva, is a type of subtropical forest found in areas with high humidity and relatively stable, mild temperatures. The forest is characterized by broadleaf tree species with evergreen, glossy and elo ...
, mostly from the tropics or warmer temperate areas, and like most of their counterparts laurifolia in the world, they are vigorous species with a great ability to populate conducive habitats. The natural habitat of most of species are in rainforest which are cloud-covered for much of the year. These species are found in forests that face threats of destruction by human
deforestation Deforestation or forest clearance is the removal of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms, ranches, or urban use. The most concentrated d ...
. Some species are in danger of extinction due to loss of habitat. ''Cryptocarya'' is a genus of great ecological importance. It is present in low rainforest and montane rainforest,
laurel forest Laurel forest, also called laurisilva or laurissilva, is a type of subtropical forest found in areas with high humidity and relatively stable, mild temperatures. The forest is characterized by broadleaf tree species with evergreen, glossy and elo ...
, in the weed-tree forests in valleys, and mixed forests of coniferous and deciduous broad-leaved trees. The differences are ecological adaptations to different environments over a relatively dry-wet and a warmer to mild frost (-2 Â°C) in temperate climate growing in cooler regions, subject to frost and occasional snow. The Chilean ''
Cryptocarya alba ''Cryptocarya alba'', the peumo or Chilean acorn, is an evergreen tree that grows in Chile and Argentina from 33 to 40° southern latitude. It can live both in wet and as in dry conditions. Its distribution can reach up to 1500 meters (5000  ...
'' and the Australian '' C. erythroxylon'' and '' C. foveolata'' of the mountains of
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
are outstanding for their frost tolerance within a genus having its majority of species growing in
tropical climate Tropical climate is the first of the five major climate groups in the Köppen climate classification identified with the letter A. Tropical climates are defined by a monthly average temperature of 18 Â°C (64.4 Â°F) or higher in the cool ...
s. Species in less-humid environments are smaller or less robust, with less abundant and thinner foliage and have oleifera cells that give trees a more fragrant aroma. The fruit, a drupe, is an important food source for birds, usually from specialized genera. Birds eat the whole fruit and regurgitate seeds intact, expanding the seeds in the best conditions for germination ( ornitochory). In some species,
seed dispersal In Spermatophyte plants, seed dispersal is the movement, spread or transport of seeds away from the parent plant. Plants have limited mobility and rely upon a variety of dispersal vectors to transport their seeds, including both abiotic vectors, ...
is carried out by
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s.


Human use

The most known trees are used by the timber industry. In this genus, the wood of some species has high commercial value. ''C. alba'', the
peumo Peumo is a Chilean city and commune in Cachapoal Province, O'Higgins Region. It is believed that Peumo was named for the vast amount of Cryptocarya alba, or Peumo trees, that grow there. Demographics According to the 2002 census of the Nation ...
, the most common evergreen tree in the
Chilean Matorral The Chilean Matorral (NT1201) is a terrestrial ecoregion of central Chile, located on the west coast of South America. It is in the Mediterranean forests, woodlands, and scrub biome, part of the Neotropical realm. Matorral is typically character ...
ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of l ...
of central Chile, produces edible reddish fruits and is the most known species in the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
. ''C. massoy'' is used commercially to produce
essential oil An essential oil is a concentrated hydrophobic liquid containing volatile (easily evaporated at normal temperatures) chemical compounds from plants. Essential oils are also known as volatile oils, ethereal oils, aetheroleum, or simply as the o ...
s. ''C. woodii'' leaves have been found in prehistoric settlements in Africa and are believed to have been used for insect control.
Essential oil An essential oil is a concentrated hydrophobic liquid containing volatile (easily evaporated at normal temperatures) chemical compounds from plants. Essential oils are also known as volatile oils, ethereal oils, aetheroleum, or simply as the o ...
is commercially harvested from '' Cryptocarya agathophylla'' (formerly ''Ravensara aromatica''), a tree native to the lowland rainforests of eastern Madagascar. Known as Ravensara oil, it is used for
aromatherapy Aromatherapy is based on the usage of aromatic materials including essential oils and other aroma compounds, with claims for improving psychological and physical well-being. It is offered as a complementary therapy or as a form of alternative m ...
in Europe and America.


Some ''Cryptocarya'' species

For full list of the 357 Cryptocarya species see List of Cryptocarya species *'' Cryptocarya agathophylla'' – tavolobelelo or clove nutmeg (Madagascar) *''
Cryptocarya alba ''Cryptocarya alba'', the peumo or Chilean acorn, is an evergreen tree that grows in Chile and Argentina from 33 to 40° southern latitude. It can live both in wet and as in dry conditions. Its distribution can reach up to 1500 meters (5000  ...
'' — peumo (Chile) *''
Cryptocarya angulata ''Cryptocarya'' is a genus of evergreen trees belonging to the laurel family, Lauraceae. The genus includes more than 350 species, distributed through the Neotropical, Afrotropical, Indomalayan, and Australasian realms. Overview The genus incl ...
'' — ivory laurel (Australia) *''
Cryptocarya ashersoniana ''Cryptocarya'' is a genus of evergreen trees belonging to the laurel family, Lauraceae. The genus includes more than 350 species, distributed through the Neotropical, Afrotropical, Indomalayan, and Australasian realms. Overview The genus incl ...
'' (Brazil) *'' Cryptocarya bidwillii'' — yellow laurel (Australia) *'' Cryptocarya caesia'' *''
Cryptocarya chinensis ''Cryptocarya chinensis'', commonly known as the Chinese cryptocarya, is a medium-sized evergreen tree native to the subtropical forests of Taiwan, southern China (Fujian, Guangdong, Guangxi, Hainan, Sichuan), and Japan Japan ( ja, æ—¥æ ...
'' *'' Cryptocarya cinnamomifolia'' (Australia) *'' Cryptocarya crassinervia'' (Indonesia) *'' Cryptocarya cunninghamiana'' — Cunningham's laurel (Australia) *'' Cryptocarya densiflora'' (Indonesia) *'' Cryptocarya elegans'' *''
Cryptocarya erythroxylon ''Cryptocarya erythroxylon'' is a medium to large rainforest tree, that grows from the Barrington Tops in New South Wales to the Gladstone area in Queensland. The common name is the pigeonberry ash, rose maple, or rose walnut. This is one o ...
'' — pigeonberry ash (southeast Australia) *'' Cryptocarya floydii'' — gorge laurel (Australia) *'' Cryptocarya foetida'' — stinking laurel (Australia) -vulnerable *''
Cryptocarya foveolata ''Cryptocarya'' is a genus of evergreen trees belonging to the laurel family, Lauraceae. The genus includes more than 350 species, distributed through the Neotropical, Afrotropical, Indomalayan, and Australasian realms. Overview The genus in ...
'' — mountain walnut (southeast Australia) *''
Cryptocarya glaucescens ''Cryptocarya glaucescens'', commonly known as jackwood, is a rainforest tree of the laurel family growing in eastern Australia. Taxonomy ''Cryptocarya glaucescens'' was one of the many species first described by Scottish botanist Robert Brown ...
'' — jackwood (Australia) *'' Cryptocarya grandis'' — cinnamon laurel (Australia) *'' Cryptocarya gregsonii'' (Lord Howe Island) *''
Cryptocarya hypospodia ''Cryptocarya'' is a genus of evergreen trees belonging to the laurel family, Lauraceae. The genus includes more than 350 species, distributed through the Neotropical, Afrotropical, Indomalayan, and Australasian realms. Overview The genus inc ...
'' (Australia) *'' Cryptocarya laevigata'' — glossy laurel (Australia) *'' Cryptocarya liebertiana'' — (
Afromontane The Afromontane regions are subregions of the Afrotropical realm, one of the Earth's eight biogeographic realms, covering the plant and animal species found in the mountains of Africa and the southern Arabian Peninsula. The Afromontane regions ...
forests in Tanzania, Mozambique, Malawi, South Africa) *'' Cryptocarya mackinnoniana'' — rusty laurel (Australia) *'' Cryptocarya mannii'' (Kaua'i) *'' Cryptocarya massoy'' (Indonesia) *'' Cryptocarya meissneriana'' (Australia) *'' Cryptocarya micrantha'' (Brazil) *'' Cryptocarya microneura'' — murrogun (Australia) *'' Cryptocarya moschata'' (Brazil) *'' Cryptocarya murrayi'' — Murray's laurel (Australia) *'' Cryptocarya natalensis'' = '' Dahlgrenodendron natalense'' (South Africa) *''
Cryptocarya nigra ''Cryptocarya'' is a genus of evergreen trees belonging to the laurel family, Lauraceae. The genus includes more than 350 species, distributed through the Neotropical, Afrotropical, Indomalayan, and Australasian realms. Overview The genus incl ...
'' (Indonesia) *'' Cryptocarya nitens'' (Indonesia) *'' Cryptocarya nova-anglica'' — mountain laurel (Australia) *''
Cryptocarya oahuensis ''Cryptocarya'' is a genus of evergreen trees belonging to the laurel family, Lauraceae. The genus includes more than 350 species, distributed through the Neotropical, Afrotropical, Indomalayan, and Australasian realms. Overview The genus incl ...
'' (O'ahu) *''
Cryptocarya obovata ''Cryptocarya obovata'' is a species of laurel growing on basaltic and fertile alluvial soils in eastern Australian rainforests. It is found from Wyong (33° S) in New South Wales to Gympie (27° S) in the state of Queensland. Extinct in the Ill ...
'' (Australia) *'' Cryptocarya onoprienkoana'' — rose maple (Queensland) *'' Cryptocarya palawanensis'' (
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
) — endangered *'' Cryptocarya pleurosperma'' — poison walnut (Australia) — toxic and corrosive sap *'' Cryptocarya pulchrinervia'' (Indonesia) *'' Cryptocarya rigida'' — rose maple (Australia) *'' Cryptocarya scortechinii'' (Indonesia) *'' Cryptocarya triplinervis'' — brown laurel (Australia) *'' Cryptocarya wightiana'' *'' Cryptocarya williwilliana'' — small-leaved laurel (Australia) *''
Cryptocarya woodii ''Cryptocarya woodii'', the Cape quince, is a shrub or small forest tree, native to southern and eastern Africa. Its Latin name commemorates John Medley Wood, a botanist in Natal. From mid summer the tree bears small, inconspicuous flowers. The ...
'' *'' Cryptocarya sheikelmudiyana'' - large (India - Western Ghats - Critically EndangeredDeepu, S. & Amitha Bachan, K.H. 2021. Cryptocarya sheikelmudiyana. The IUCN Red List of Threatened Species 2021: e.T187622842A187623277. https://dx.doi.org/10.2305/IUCN.UK.2021-2.RLTS.T187622842A187623277.en) ''Cryptocarya palmerstonii is now a synonym of ''
Endiandra palmerstonii ''Endiandra palmerstonii'', popularly known as Queensland walnut or black walnut, is a rainforest tree of northern Queensland. It was named after the Australian prospector Christie Palmerston. Queensland walnut has been used as a furniture timbe ...
'', Queensland walnut (Queensland).


References


External links

* Pictures o
''Cryptocarya alba'' or Peumo
in Chile. * {{Taxonbar, from=Q3309395 Lauraceae genera Taxa named by Robert Brown (botanist, born 1773)