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: ''Common names: Middle American rattlesnake, Central American rattlesnake,''Klauber LM. 1997. Rattlesnakes: Their Habitats, Life Histories, and Influence on Mankind. Second Edition. 2 volumes. Reprint, University of California Press, Berkeley. . ''tzabcan'' (local name for subspecies ''C. s. tzabcan'') ''Crotalus simus'' is a
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
pit viper The Crotalinae, commonly known as pit vipers,Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . crotaline snakes (from grc, κρόταλον ''krotalon'' castanet), or pit adders, are a subfa ...
species found in Mexico and Central America. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
is Latin for "flat-nosed", likely because its head is blunt compared with lanceheads (''
Bothrops ''Bothrops'' is a genus of highly venomous pit vipers endemic to Central and South America.McDiarmid RW, Campbell JA, Touré T. 1999. ''Snake Species of the World: A Taxonomic and Geographic Reference, vol. 1''. Herpetologists' League. 511 pp. ...
''). Three
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognized, including the nominate subspecies described here.


Description

Adults commonly exceed in length, with males growing larger than females. Large males reach in some populations. The maximum length is . The body has a rough appearance because the dorsal scale keels are accentuated into protuberances or tuberculations. This is most apparent on the scale rows on either side of the body with a decreasing intensity in the lower rows. The vertebral scales are about as prominently keeled as the fourth row down on the flanks (with the vertebral scales as the first row).


Distribution and habitat

The species is found from
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
in southwestern
Michoacán Michoacán, formally Michoacán de Ocampo (; Purépecha: ), officially the Free and Sovereign State of Michoacán de Ocampo ( es, Estado Libre y Soberano de Michoacán de Ocampo), is one of the 32 states which comprise the Federal Entities of ...
on the Pacific coast, and
Veracruz Veracruz (), formally Veracruz de Ignacio de la Llave (), officially the Free and Sovereign State of Veracruz de Ignacio de la Llave ( es, Estado Libre y Soberano de Veracruz de Ignacio de la Llave), is one of the 31 states which, along with Me ...
and the Yucatan Peninsula on the Atlantic coast, south through
Belize Belize (; bzj, Bileez) is a Caribbean and Central American country on the northeastern coast of Central America. It is bordered by Mexico to the north, the Caribbean Sea to the east, and Guatemala to the west and south. It also shares a wate ...
,
Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by H ...
,
El Salvador El Salvador (; , meaning " The Saviour"), officially the Republic of El Salvador ( es, República de El Salvador), is a country in Central America. It is bordered on the northeast by Honduras, on the northwest by Guatemala, and on the south b ...
,
Honduras Honduras, officially the Republic of Honduras, is a country in Central America. The republic of Honduras is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Oce ...
, and
Nicaragua Nicaragua (; ), officially the Republic of Nicaragua (), is the largest country in Central America, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west. Managua is the cou ...
to west-central Costa Rica. It is absent from
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
, but apparently does occur on the Atlantic side of
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
. The type locality given is "Ceylan", which is incorrect. Its habitats are semiarid, including dry or very dry tropical forests, thorn woodland, and arid scrub forest. It also occurs in
mesic forest In ecology, a mesic habitat is a type of habitat with a moderate or well-balanced supply of moisture, e.g., a mesic forest, a temperate hardwood forest, or dry-mesic prairie. Mesic habitats transition to xeric shrublands in a non-linear fashion, ...
s with relatively dry, open areas.


Uses

To the
Mayans The Maya peoples () are an ethnolinguistic group of indigenous peoples of Mesoamerica. The ancient Maya civilization was formed by members of this group, and today's Maya are generally descended from people who lived within that historical reg ...
, the Yucatan subspecies (''C. s. tzabcan'') is greatly revered. The word ''tzabcan'' means rattlesnake in Mayan. What the rattlesnake actually symbolizes is unknown, but many temples have carved rattlesnake shapes. Shamans also dry and roast snakes, grinding them into a powder used as medicine.


Venom

Bites are similar to
rattlesnake Rattlesnakes are venomous snakes that form the genera ''Crotalus'' and ''Sistrurus'' of the subfamily Crotalinae (the pit vipers). All rattlesnakes are vipers. Rattlesnakes are predators that live in a wide array of habitats, hunting small anim ...
bites in the United States. Local symptoms may be severe, with pain, massive swelling, blistering, and necrosis that lead to fasciotomies and in some cases
amputation Amputation is the removal of a limb by trauma, medical illness, or surgery. As a surgical measure, it is used to control pain or a disease process in the affected limb, such as malignancy or gangrene. In some cases, it is carried out on indi ...
s. Systemic effects involving hemostatic disturbances are rare, as are
kidney failure Kidney failure, also known as end-stage kidney disease, is a medical condition in which the kidneys can no longer adequately filter waste products from the blood, functioning at less than 15% of normal levels. Kidney failure is classified as eit ...
, and neurotoxicity. Only venom from neonates contains crotoxin; a constituent typically found in '' C. durissus'' venom that produces neurotoxic symptoms.Warrell DA. 2004. Snakebites in Central and South America: Epidemiology, Clinical Features, and Clinical Management. ''In'' Campbell JA, Lamar WW. 2004. The Venomous Reptiles of the Western Hemisphere. Comstock Publishing Associates, Ithaca and London. 870 pp. 1500 plates. .


Taxonomy

Previously, until 2004, the description for this form was listed as the nominate subspecies for the tropical rattlesnake, '' C. durissus''. Molecular genetic data suggest the taxa ''culminatus'' and ''tzabcan'' should be considered as separate species from ''C. simus''Wüster, W., J.E. Ferguson, J.A. Quijada-Mascareñas, C.E. Pook, M.G. Salomão & R.S. Thorpe (2005) Tracing an invasion: landbridges, refugia and the phylogeography of the Neotropical rattlesnake (Serpentes: Viperidae: Crotalus durissus). Molecular Ecology 14(4): 1095–1108.


References


Further reading

* Sonnini, C.S. & Latreille, P.A. 1801. ''Histoire naturelle des Reptiles, avec figures dessinées d'après nature; Tome III. Seconde Partie. Serpens.'' Crapelet. Paris. 335 pp. (''Crotalus simus'', p. 202.) {{Taxonbar, from=Q146338 simus Reptiles of Guatemala Reptiles described in 1801