Croesia Semipurpurana
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''Acleris semipurpurana'' is a species in the moth family Tortricidae, and one of several species of moth commonly known as oak leaftier or oak leaf tier. The larvae feed on the leaves of oak trees in the Eastern United States and southeastern Canada which can be a major cause of defoliation. The loss of leaves can kill or damage the affected trees, which are chiefly in the ''Lobatae'' or red oak section of '' Quercus'', or oaks. Adult ''Acleris semipurpurana'' moths lay single eggs on oak tree branches in June which develop over winter and hatch the next April. The larvae eat tree buds and young leaves, then tie leaves together with silk (hence the name). They nest and eat inside the tied leaves, then pupate in the leaf litter on the floor in May or June. After a week or two the adult moths emerge, mate, and lay the next generation of eggs.


Life cycle and description

Oak leaftier moths lay their eggs in June each year, typically as single eggs on rough-bark twigs, typically "second year branches". Note: this is published on the ForestPests.org website with the title "Oak Leaftiers: ''Acleris semipurpurana'' (Kearfott)" The eggs overwinter, then hatch in the following spring. After larvae (or
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s) emerge in April, they eat the buds of oak trees and the young leaves inside them. When they are fully grown, the larvae are long and "dirty white to light green" with "a pale brown head and black to brown front legs". Note: this is published on the ForestPests.org website with the title "Oak leaftier: ''Acleris semipurpurana''" Other identifying characteristics of the larvae are a long head with a black bar on the cheek or lower portion, and brown legs on the thorax. The larvae tie leaves together with silk, which gives the insect its common name. The larvae feed and nest inside the tied or folded leaves until mid- to late May, when they begin to pupate. The larvae leave the trees to pupate, descending to the ground on silken threads. The pupae are commonly found in leaf litter on the forest floor beneath the oak trees. In late May or June, after one or two weeks in the pupal stage, adult moths emerge. The moths are small with a wingspan of ; their wings have a characteristic bell shape. Wing colors can vary considerably. Some forewings are yellow while others have dark brown or even purple markings. The adults mate and lay eggs - only one generation of moths is produced annually. ''Acleris semipurpurana'' was first described by
William D. Kearfott William D. Kearfott (January 12, 1864 – November 12, 1917) was an American engineer and amateur entomologist. Kearfott was educated in primary schools in Richmond and Philadelphia and was connected with the Morton Poole Company and the Internati ...
in 1909.


Range and impact

''Acleris semipurpurana'' is found in the eastern United States and adjoining portions of southeastern Canada. It has been found in US states ranging from
New York New York most commonly refers to: * New York City, the most populous city in the United States, located in the state of New York * New York (state), a state in the northeastern United States New York may also refer to: Film and television * '' ...
, Massachusetts, Connecticut and Pennsylvania in the northeast to Minnesota and Texas in the west, as well as the Canadian province of Ontario. The oak leaftiers feed mainly trees in the ''Lobatae'' or red oak section of '' Quercus'' or oak trees: Specifically on
eastern black oak ''Quercus velutina'', the black oak, is a species of oak in the red oak group (''Quercus'' sect. ''Lobatae''), native and widespread in eastern and central North America. It is sometimes called the eastern black oak. ''Quercus velutina'' was pre ...
, northern red oak, pin oak,
scarlet oak ''Quercus coccinea'', the scarlet oak, is a deciduous tree in the red oak section ''Lobatae'' of the genus ''Quercus'', in the family Fagaceae. It is primarily distributed in the central and eastern United States. It occurs on dry, sandy, usuall ...
, and
scrub oak Scrub oak is a common name for several species of small, shrubby oaks. It may refer to: *the Chaparral plant community in California, or to one of the following species. In California *California scrub oak (''Quercus berberidifolia''), a widespr ...
. Several species of moths are referred to with the common name oak leaftier, most from the family Tortricidae, but also from "three other minor families". Of these species ''Acleris semipurpurana'' is the most common and one of the most important for defoliation. In addition to the oak leaftiers, other pest species will often feed on the same trees, including ''
Archips semiferanus ''Archips semiferanus'' (also known as ''Archips semiferana'') is a species of moth in the family Tortricidae, and one of several species of moth commonly known as oak leafroller or oak leaf roller. The larvae feed on the leaves of oak trees in ...
'' and other oak leafrollers. The young larvae eat the buds and either destroy developing leaves, or cause leaves to develop with many holes in them. This can severely stress the tree's food reserves, and the older larvae can eat nearly all the remaining leaves, defoliating them. When trees are defoliated two or more years in a row, this leads to "branch dieback, loss of diameter growth, and tree decline". The oak leaftier has caused "considerable tree mortality", especially in the Appalachian region. Dead wood in affected trees is attacked by fungi such as the shoestring root fungus and wood borers like the twolined chestnut borer. A major outbreak occurred in the 1960s and 1970s in Connecticut, Massachusetts, New York, Pennsylvania, Virginia, and West Virginia. Tiadaghton State Forest in north central Pennsylvania was especially hard hit. Natural methods of pest control usually are sufficient to keep ''Croesia semipurpurana'' populations in check, but chemical
pesticide Pesticides are substances that are meant to control pests. This includes herbicide, insecticide, nematicide, molluscicide, piscicide, avicide, rodenticide, bactericide, insect repellent, animal repellent, microbicide, fungicide, and lampri ...
s are sometimes needed.


References


External links


Forestryimages.org: photos of ''Acleris semipurpurana''
— ''synonym: Croesia semipurpurana''. {{Taxonbar , from=Q4674314 semipurpurana Moths of North America Moths described in 1909