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''Aquilegia formosa'', the crimson columbine, western columbine, or (ambiguously) "red columbine", is a common
wildflower A wildflower (or wild flower) is a flower that grows in the wild, meaning it was not intentionally seeded or planted. The term implies that the plant probably is neither a hybrid nor a selected cultivar that is in any way different from the w ...
native to western
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, from
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
to
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
, and eastward to
Montana Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...
and
Wyoming Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the south ...
.


Description

''Aquilegia formosa'' is a perennial herb that grows to 20–80 cm in height, averaging around 60 cm. Flowers, which can be seen from April to August (with some variation between regions), are about 5 cm long and red and yellow in color. The sepals and petal spurs are typically a reddish-orange color, attributed to the anthocyanin pigments pelargonidin and cyanidin, and carotenoids. Petal blades are yellow, pigmented by carotenoids. The primary pollinators are hummingbirds, although bees, butterflies, and flies will also visit flowers. Despite several floral adaptations to hummingbird pollination, at ~9,000-10,000 feet in elevation in the eastern drainages of the central Sierra Nevada mountains of California, ''A. formosa'' forms hybrid zones with ''Aquilegia pubescens'', which is primarily pollinated by hawk moths. The flowers are edible, with a sweet taste—though the seeds can be fatal if eaten, and most parts of the plant contain
cyanogenic glycoside In chemistry, a glycoside is a molecule in which a sugar is bound to another functional group via a glycosidic bond. Glycosides play numerous important roles in living organisms. Many plants store chemicals in the form of inactive glycosides. ...
s.


Distribution

Within its range, the crimson columbine can be found in most kinds of habitat (
chaparral Chaparral ( ) is a shrubland plant community and geographical feature found primarily in the U.S. state of California, in southern Oregon, and in the northern portion of the Baja California Peninsula in Mexico. It is shaped by a Mediterranean c ...
,
oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...
woodland, mixed-evergreen or
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
ous forest). It is not found on desert floors, nor at altitudes above 3300 metres, and it is absent from the Central Valley of
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
. It prefers moist locations such as stream banks.


Native American use

Some
Plateau Indian Indigenous peoples of the Northwest Plateau, also referred to by the phrase Indigenous peoples of the Plateau, and historically called the Plateau Indians (though comprising many groups) are indigenous peoples of the Interior of British Columbia ...
tribes used the ''Aquilegia formosa'' to concoct a perfume. It is also used medicinally by several Native American tribes.


Etymology

''Aquilegia'' is derived from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
word 'aquila', meaning 'eagle', or possibly from the
Medieval In the history of Europe, the Middle Ages or medieval period lasted approximately from the late 5th to the late 15th centuries, similar to the Post-classical, post-classical period of World history (field), global history. It began with t ...
German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ger ...
words 'Acheleia' or 'Akelei'; this name is in reference to its talon-like nectaries. ''Formosa'' means 'handsome', 'beautiful', or 'well-formed'.Gledhill, David (2008). "The Names of Plants". Cambridge University Press. (hardback), (paperback). pp 53, 169


References


External links


Calflora Database: ''Aquilegia formosa'' (western columbine, crimson columbine)

Jepson Manual eFlora (TJM2) treatment of ''Aquilegia formosa''

UC Photos gallery — ''Aquilegia formosa''
{{Taxonbar, from=Q1093527
formosa Taiwan, officially the Republic of China (ROC), is an island country located in East Asia. The main island of Taiwan, formerly known in the Western political circles, press and literature as Formosa, makes up 99% of the land area of the territorie ...
Flora of the Northwestern United States Flora of Western Canada Flora of Alaska Flora of Baja California Flora of California Flora of the Great Basin Flora of the Rocky Mountains Flora of the Sierra Nevada (United States) Natural history of the California chaparral and woodlands Plants used in traditional Native American medicine Garden plants of North America Butterfly food plants Flora without expected TNC conservation status