The inland riffle sculpin (''Cottus gulosus'') is a species of

fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

in the family

Cottidae The Cottidae are a family of fish in the superfamily Cottoidea, the sculpins. It is the largest sculpin family, with about 275 species in 70 genera.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology ...

. It is found in the

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

, inhabiting the lower

Columbia River The Columbia River (Upper Chinook: ' or '; Sahaptin: ''Nch’i-Wàna'' or ''Nchi wana''; Sinixt dialect'' '') is the largest river in the Pacific Northwest region of North America. The river rises in the Rocky Mountains of British Columbia, C ...

drainage in

Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered on ...

, to Morro Bay in

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

. It is also found in the

Puget Sound Puget Sound ( ) is a sound of the Pacific Northwest, an inlet of the Pacific Ocean, and part of the Salish Sea. It is located along the northwestern coast of the U.S. state of Washington. It is a complex estuarine system of interconnected ma ...

drainage in Washington. It reaches a maximum length of 11.0 cm. It prefers rocky riffles of headwaters and creeks.

Taxonomy

The inland riffle sculpin was first formally described as ''Cottopsis gulosus'' by the French biologist

Charles Frédéric Girard Charles Frédéric Girard (8 March 1822 – 29 January 1895) was a French biologist specializing in ichthyology and herpetology. Born in Mulhouse, France, he studied at the College of Neuchâtel, Switzerland, as a student of Louis Agassiz. In ...

with its type locality given as the San Joaquin River in California. This species is classified by some authorities in the

subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...

''Cottopsis''.

Description

Inland riffle sculpins have a brown/tan coloration mottled with dark brown spots and reddish hues around their fins. They have large mouths with a single median chin pore, and often times will feature joined dorsal fins. Despite not being considered particularly definitive in their appearance, riffle sculpins can still be identified through notes regarding specific spotting and ray numbers. In addition to their mottled color, riffle sculpins have a large black spot near the end of the first dorsal fin. They have 16-19 rays in their second dorsal fin, 15-16 rays in each of their pectoral fins, and 12-16 rays in their anal fin. Spawning male riffle sculpins can be seen to have a darker appearance coupled with an orange hue at the edge of the first dorsal fin.

Habitat

Inland riffle sculpins can often be found in headwater streams with riffles. When cohabiting alongside prickly sculpin, riffle sculpin choose to move towards the cooler areas of the stream. However, they are also found to have a preference for permanent streams with stable temperatures of 25-26 °C. When available, riffle sculpins will move towards areas of cover, such as rocks or overhanging banks. Riffles act as preferred habitats to multiple species due to the many prey organisms that live in the area. However, the division of habitat within riffle sculpin and other species such as the speckled dace is driven primarily by temperature, as the metabolic rates of the sculpin rapidly increase alongside it.

Diet

Inland riffle sculpins maintain a diet primarily built of benthic invertebrates and active insect larvae. This includes insects like mayflies and caddisflies. If easily available, riffle sculpins will prey on small fish and amphipods, typically feeding during the night.

References

{{Taxonbar, from=Q5630226 Cottus (fish) Fish described in 1854 Taxa named by Charles Frédéric Girard