Cosmopterix Pulchrimella
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''Cosmopterix pulchrimella'', the beautiful cosmopterix moth, is a
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
of the family
Cosmopterigidae The Cosmopterigidae are a family of insects (cosmet moths) in the order Lepidoptera. These are small moths with narrow wings whose tiny larvae feed internally on the leaves, seeds and stems of their host plants. About 1500 species are described. ...
. It is known from the United States (from
Massachusetts Massachusetts (Massachusett language, Massachusett: ''Muhsachuweesut assachusett writing systems, məhswatʃəwiːsət'' English: , ), officially the Commonwealth of Massachusetts, is the most populous U.S. state, state in the New England ...
,
Michigan Michigan () is a state in the Great Lakes region of the upper Midwestern United States. With a population of nearly 10.12 million and an area of nearly , Michigan is the 10th-largest state by population, the 11th-largest by area, and the ...
, and southern
Wyoming Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the south ...
south to southern
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
, southern
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
, and
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
) and
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
. It is also present in the
Palearctic realm The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
, where it is known from the
Mediterranean Basin In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and w ...
, from
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
to the western
Transcaucasus The South Caucasus, also known as Transcaucasia or the Transcaucasus, is a geographical region on the border of Eastern Europe and Western Asia, straddling the southern Caucasus Mountains. The South Caucasus roughly corresponds to modern Arme ...
, north to
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
and
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia a ...
. It has also been recorded from the
Azores ) , motto =( en, "Rather die free than subjected in peace") , anthem= ( en, "Anthem of the Azores") , image_map=Locator_map_of_Azores_in_EU.svg , map_alt=Location of the Azores within the European Union , map_caption=Location of the Azores wi ...
, the
Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...
and
Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...
. It has recently been found in southern
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
.


Description


Adult

Male, female. Forewing length 3.0-4.0 mm. Head: frons shining grey, shining white towards clypeus, vertex and neck tufts shining dark brown, laterally and medially lined white, collar shining dark brown; labial palpus first segment very short, white, second segment four-fifths of the length of third, shining white on inside, dark brown with a white longitudinal line on outside, third segment white, lined brown laterally; scape dorsally shining dark brown with a white anterior line, ventrally shining white, antenna shining dark brown with a white interrupted line from base to beyond one-half, near base often uninterrupted, followed towards apex by five dark brown segments, one white, one dark brown, one white, three dark brown, one white, approximately twelve dark brown, three white and five dark brown segments at apex. Thorax and tegulae shining dark brown, thorax with a white median line. Legs: shining dark brown, foreleg with a white line on tibia and tarsal segment one, tibia of midleg with white oblique basal and medial lines and a white apical ring, tarsal segments one and two with white apical rings, segment five entirely white, tibia of hindleg as midleg, tarsal segments four and five entirely white, spurs white dorsally, brown ventrally. Forewing shining dark brown, three short silver streaks in the basal area, sometimes with a pale golden gloss, a subcostal nearest to base and bending from costa distally, a shorter medial underneath the distal end of the subcostal, a subdorsal, as long as the subcostal but further from base, a bright orange transverse fascia beyond the middle, narrowing towards dorsum, in some of the specimens from the USA the fascia is partly or completely darkened, bordered at the inner edge by broad tubercular pale golden metallic fascia, sometimes with a pinkish gloss, perpendicular at dorsum, edged blackish on the outside, bordered at the outer edge by a similarly coloured inwardly oblique fascia, edged blackish on the inside, at the costa of the outer fascia a broad white costal streak, the apical line as a silver metallic spot with bluish reflection in the middle of the apical area and a broad white spot in the cilia at apex, cilia dark brown. Hindwing shining brown, cilia brown. Underside: forewing shining dark greyish brown, the white costal streak and apical spot distinctly visible, hindwing shining dark greyish brown. Abdomen shining dark brown dorsally, ventrally segments banded white posteriorly, anal tuft brown.The genera Cosmopterix Hübner and Pebobs Hodges in the New World with special attention to the Neotropical fauna (Lepidoptera: Cosmopterigidae)
/ref>


Larva

Head and prothoracic plate black, anal plate greyish brown, thoracic legs pale brown, body greyish white.


Biology

The larvae feed on ''
Parietaria judaica ''Parietaria judaica'', with common names spreading pellitory or pellitory of the wall, is a species of herbaceous perennial plant in the family Urticaceae. The plant's pollen is highly allergenic. In Australia it is also known as asthma weed, du ...
'', ''
Parietaria officinalis ''Parietaria officinalis'', the eastern pellitory-of-the-wall, also known as upright pellitory and lichwort, is a plant of the nettle family. Its leaves, however, are non-stinging. The plant grows on rubbish and on walls, hence the name. The pol ...
'', ''
Parietaria pensylvanica ''Parietaria pensylvanica'', commonly called Pennsylvania pellitory, is a species of flowering plant in the nettle family. It is native to much of North America including every province in Canada except the Maritimes and Newfoundland and Labrador ...
'' and ''
Pilea pumila ''Pilea pumila'', commonly known as clearweed, Canadian clearweed, coolwort or richweed, is an herbaceous plant in the nettle family (Urticaceae). It is native to Asia and eastern North America, where it is broadly distributed. This plant is mos ...
''. They
mine Mine, mines, miners or mining may refer to: Extraction or digging * Miner, a person engaged in mining or digging *Mining, extraction of mineral resources from the ground through a mine Grammar *Mine, a first-person English possessive pronoun ...
the leaves of their host plant. The mines starts as an irregular gallery usually at the midrib, soon leading to an irregular blotch. Inside the mine a silken spinning is found, which often causes contortion of the leaf. The larva constructs a silk-lined gallery, which serves as a shelter when the larva is not feeding. The
frass Frass refers loosely to the more or less solid excreta of insects, and to certain other related matter. Definition and etymology ''Frass'' is an informal term and accordingly it is variously used and variously defined. It is derived from the ...
is partly inside the mine, but most of it is rejected through a hole at the beginning of the mine. The black pellets of frass on the underside of a leaf indicate the presence of a larva. New mines are made very often. The cocoon is found inside the mine. The species is bivoltine, with adults on wing throughout the year in southern areas.


References


External links


''Cosmopterix pulchrimella'' at UKmoths
{{Taxonbar, from=Q291678 pulchrimella Leaf miners Moths described in 1875 Moths of North America Moths of Europe