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The coscoroba swan (''Coscoroba coscoroba'') is a species of
waterfowl Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which in ...
in subfamily
Anserinae The Anserinae are a subfamily in the waterfowl family Anatidae. It includes the swans and true geese. Under alternative systematical concepts (see e.g., Terres & NAS, 1991), it is split into two subfamilies, the Anserinae contain the geese and ...
of the family
Anatidae The Anatidae are the biological family of water birds that includes ducks, geese, and swans. The family has a cosmopolitan distribution, occurring on all the world's continents except Antarctica. These birds are adapted for swimming, floating ...
.HBW and BirdLife International (2021) Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world. Version 6. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v6_Dec21.zip retrieved August 7, 2022 It is found in
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
,
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
,
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
,
Paraguay Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to th ...
,
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
, and the
Falkland Islands The Falkland Islands (; es, Islas Malvinas, link=no ) is an archipelago in the South Atlantic Ocean on the Patagonian Shelf. The principal islands are about east of South America's southern Patagonian coast and about from Cape Dubouzet ...
.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 July 2022. Species Lists of Birds for South American Countries and Territories. https://www.museum.lsu.edu/~Remsen/SACCCountryLists.htm retrieved July 24, 2022


Taxonomy and systematics

The coscoroba swan's placement within the family Anatidae is not settled. A 2014 genetic study posited a phylogenetic relationship between this species and the
Cape Barren goose The Cape Barren goose (''Cereopsis novaehollandiae'') is a large goose resident in southern Australia. Etymology The species' common name is derived from Cape Barren Island, where specimens were first sighted by European explorers. It is known ...
(''Cereopsis novaehollandiae'').
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
's ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. T ...
'' has adopted this approach and places the coscoroba swan and Cape Barren goose together in tribe Cereopsini, which it places as basal to ''Cygnus'' and all other geese. However, the South American Classification Committee of the
American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its m ...
, the
International Ornithological Committee The International Ornithologists' Union, formerly known as the International Ornithological Committee, is a group of about 200 international ornithologists, and is responsible for the International Ornithological Congress and other international ...
(IOC), and the
Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 2022 ...
treat the Cape Barren goose as basal to other geese, which themselves precede the coscoroba and other swans in a linear sequence. Even those three systems differ, with the IOC placing the coscoroba before ''Cygnus'' and the other two after it in their lists.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 July 2022. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved July 24, 2022Clements, J. F., T. S. Schulenberg, M. J. Iliff, S. M. Billerman, T. A. Fredericks, J. A. Gerbracht, D. Lepage, B. L. Sullivan, and C. L. Wood. 2021. The eBird/Clements checklist of Birds of the World: v2021. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ Retrieved August 25, 2021 The coscoroba swan is the only member of genus ''Coscoroba'' and has no subspecies.


Description

The coscoroba is the smallest swan at long with a wingspan of about . Males weigh and females . Their appearance is intermediate between those of geese and other swans. The sexes have the same plumage; adults are white except for black tips to the outer six
primary feather Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...
s. Although this black is often barely visible on the closed wing it is conspicuous in flight. Their beak, legs, and feet are red. The
cygnet A cygnet is a young swan. Cygnet may also refer to: Places *Cygnet Island, a small islet in south-eastern Australia *Cygnet, Ohio, a village in the United States *Cygnet River, South Australia, a locality on Kangaroo Island *Cygnet, Tasmania, a ...
is grayish white to brownish with a black cap and a blue-gray bill, and attains adult plumage at about eight months of age.Carboneras, C. and G. M. Kirwan (2020). Coscoroba Swan (''Coscoroba coscoroba''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.cosswa1.01 retrieved September 28, 2022


Distribution and habitat

The coscoroba swan is a year-round resident of central Argentina and along the Uruguay-southern Brazil coast. It also breeds but does not winter from southern Chile and Argentina south to
Tierra del Fuego Tierra del Fuego (, ; Spanish for "Land of the Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South American mainland, across the Strait of Magellan. The archipelago consists of the main island, Isla G ...
and occasionally on the Falkland Islands. In winter its range extends north to central Chile, northern Argentina, Paraguay, Uruguay, and southern Brazil. It has been recorded as a
vagrant Vagrancy is the condition of homelessness without regular employment or income. Vagrants (also known as bums, vagabonds, rogues, tramps or drifters) usually live in poverty and support themselves by begging, scavenging, petty theft, temporar ...
in Bolivia and at several locations in Brazil north of its usual limit. The coscoroba swan inhabits well-vegetated lagoons and freshwater swamps and sometimes human-made reservoirs. It is mainly a bird of the lowlands though there are scattered records as high as and at least one at .


Behavior


Feeding

The coscoroba swan's diet has not been studied in detail, but it apparently feeds on aquatic and some terrestrial plants, small aquatic invertebrates, and small fish. It forages mainly while swimming or wading in shallow water and in contrast to other swans seldom upends. It also grazes on land. It often feeds with
black-necked swan The black-necked swan (''Cygnus melancoryphus'') is a species of waterfowl in tribe Cygnini of subfamily Anserinae.HBW and BirdLife International (2021) Handbook of the Birds of the World and BirdLife International digital checklist of the bird ...
s (''Cygnus melanocoryphus'').


Breeding

The coscoroba swan breeds in the local spring, which is May to October in northern Argentina, July to December in Chile, and July and August in southern Brazil. The species forms long-term pair bonds. Its nest is a mound of vegetation lined with soft grass constructed by both members of a pair on a small islet, partially floating in a reedbed, or in long grass close to water. The clutch size is four to nine oval eggs that weigh about with dimensions of about . Males guard females during the incubation period of about 35 days. Both parents care for the young through fledging at 14 weeks and beyond, sometimes until they are a year old. In captivity coscoroba swans can live 20 years.


Vocalization

The coscoroba swan makes an
onomatopoeic Onomatopoeia is the process of creating a word that phonetically imitates, resembles, or suggests the sound that it describes. Such a word itself is also called an onomatopoeia. Common onomatopoeias include animal noises such as ''oink'', ''m ...
"''cos-cor-oo''", usually as a threat to intruders. They also make a "monosyllabic hooting note" as a contact call between mates. Immature birds make "loud chirps and trills".


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has assessed the coscoroba swan as being of Least Concern. It has a large range and its estimated population of 6700 to 17,000 mature individuals is believed to be stable. No immediate threats have been identified. The species' population varies dramatically at different parts of its range, being fairly common in Argentina, uncommon in Paraguay and Uruguay, rare in Chile, and uncertain in Brazil. The " eatest threat seems to be loss of temperate marsh habitats due to urbanization and agricultural developments".


Gallery

File:Coscoroba coscoroba -Mostardas, Rio Grande do Sul, Brasil -juvenile-8.jpg, Juveniles in
Rio Grande do Sul Rio Grande do Sul (, , ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative_units_of_Brazil#List, fifth-most-populous state and the List of Brazilian st ...
, Brazil File:Coscoroba coscoroba (Coscoroba Swan - Koskorobaschwan) - Weltvogelpark 2012-01.jpg, Bath taking at
Weltvogelpark Walsrode Weltvogelpark Walsrode, known as Walsrode Bird Park or Jubs in English until 2010, is a bird park located in the middle of the Lüneburg Heath in North Germany within the municipality of Bomlitz near Walsrode in the state of Lower Saxony, Germany. ...
File:Coscoroba.swan.2.arp.750pix.jpg, Swimming File:Coscoroba Swan - Berlin Zoo.jpg, At
Berlin Zoo The Berlin Zoological Garden (german: link=no, Zoologischer Garten Berlin) is the oldest surviving and best-known zoo in Germany. Opened in 1844, it covers and is located in Berlin's Tiergarten. With about 1,380 different species and over 20,2 ...
, Germany


References


External links

* {{Taxonbar, from=Q144218
coscoroba swan The coscoroba swan (''Coscoroba coscoroba'') is a species of waterfowl in subfamily Anserinae of the family Anatidae.HBW and BirdLife International (2021) Handbook of the Birds of the World and BirdLife International digital checklist of the bi ...
coscoroba swan The coscoroba swan (''Coscoroba coscoroba'') is a species of waterfowl in subfamily Anserinae of the family Anatidae.HBW and BirdLife International (2021) Handbook of the Birds of the World and BirdLife International digital checklist of the bi ...
Birds of Argentina Birds of the Falkland Islands
coscoroba swan The coscoroba swan (''Coscoroba coscoroba'') is a species of waterfowl in subfamily Anserinae of the family Anatidae.HBW and BirdLife International (2021) Handbook of the Birds of the World and BirdLife International digital checklist of the bi ...