Corydalidae
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The family Corydalidae contains the megalopterous
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s known as
dobsonflies Dobsonflies are a subfamily of insects, Corydalinae, part of the Megalopteran family Corydalidae. The larvae (commonly called hellgrammites) are aquatic, living in streams, and the adults are often found along streams as well. The nine genera o ...
and
fishflies Fishflies are members of the subfamily Chauliodinae, belonging to the megalopteran family (biology), family Corydalidae. They are most easily distinguished from their closest relatives, Dobsonfly, dobsonflies, by the jaws (mandibles) and antenna ...
. Making up about one dozen
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
,See references in Haaramo (2008) they occur primarily throughout the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
, both temperate and tropical, and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
. They are sizeable Megaloptera, with a body usually larger than 25 mm (1 inch). They often have long filamentous antennae, though in male fishflies they are characteristically feathered.
Ocelli A simple eye (sometimes called a pigment pit) refers to a form of eye or an optical arrangement composed of a single lens and without an elaborate retina such as occurs in most vertebrates. In this sense "simple eye" is distinct from a multi-l ...
are present; the fourth tarsal segment is
cylinder A cylinder (from ) has traditionally been a three-dimensional solid, one of the most basic of curvilinear geometric shapes. In elementary geometry, it is considered a prism with a circle as its base. A cylinder may also be defined as an infin ...
-shaped. The four large
wings A wing is a type of fin that produces lift while moving through air or some other fluid. Accordingly, wings have streamlined cross-sections that are subject to aerodynamic forces and act as airfoils. A wing's aerodynamic efficiency is expresse ...
are translucent, smoky grey, or mixed, and the
anterior Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
pair is slightly longer than the posterior one. Their aquatic larvae are used as fish bait and are called hellgrammites. The eastern dobsonfly, ''
Corydalus cornutus The eastern dobsonfly, ''Corydalus cornutus'', is a large insect in the Corydalidae family. It is found in eastern North America in regions with fast-flowing streams where its aquatic larvae develop. These are known as hellgrammites and are amon ...
'', is the most well-known
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
n
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
among the dobsonflies. These
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
have distinctive elongated mandibles in males and form the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Corydalinae Dobsonflies are a subfamily of insects, Corydalinae, part of the Megalopteran family Corydalidae. The larvae (commonly called hellgrammites) are aquatic, living in streams, and the adults are often found along streams as well. The nine genera o ...
. The genera in which the males have normal mandibles, called fishflies, form the subfamily Chauliodinae. The summer fishfly, ''
Chauliodes pectinicornis ''Chauliodes pectinicornis'' known as Summer fishfly, is a species of fishfly from North America. Distribution ''C. pectinicornis'' has a wide distribution in the eastern Canada and United States, from Maine in the north-east to Alachua, Lib ...
'', is perhaps the best-known of these in North America; its immense mating swarms in the
Upper Mississippi River The Upper Mississippi River is the portion of the Mississippi River upstream of St. Louis, Missouri, United States, at the confluence of its main tributary, the Missouri River. History In terms of geologic and hydrographic history, the Upper ...
region fill the air on a few summer nights each year much like
mayflies Mayflies (also known as shadflies or fishflies in Canada and the upper Midwestern United States, as Canadian soldiers in the American Great Lakes region, and as up-winged flies in the United Kingdom) are aquatic insects belonging to the orde ...
in certain regions of
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
, leaving millions of carcasses to be cleaned up the next day. The
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e are aquatic, active, armed with strong sharp mandibles, and breathe by means of abdominal branchial filaments. When full sized — which can take several years — they leave the water and spend a quiescent
pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...
l stage on the land, in chambers dug under stones or logs, before
metamorphosis Metamorphosis is a biological process by which an animal physically develops including birth or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and differentiation. Some inse ...
into the sexually mature insect. File:Chauliodes rastricornis P1400671a.jpg, ''Chauliodes pectinicornis'' File:Parachauliodes japonicus 01 larva.JPG, ''Parachauliodes japonicus'' File:Corydalis larva Packard.jpg, Larva


Genera

These 36 genera belong to the family Corydalidae: * '' Acanthacorydalis'' c g * '' Anachauliodes'' c g * '' Apochauliodes'' c g * ''
Archichauliodes ''Archichauliodes'' is a genus of fishflies in the family Corydalidae The family Corydalidae contains the Megaloptera, megalopterous insects known as dobsonflies and fishflies. Making up about one dozen genera,See references in Haaramo (2008) ...
'' c g * ''
Chauliodes ''Chauliodes'' is a genus of fishflies in the family Corydalidae. There are about five described species in ''Chauliodes''. Species These five species belong to the genus ''Chauliodes'': * '' Chauliodes carsteni'' Wichard, 2003 * ''Chauliodes p ...
'' Latreille, 1796 i c g b – fishflies * '' Chauliosialis'' c g * '' Chloronia'' Banks, 1908 i c g * '' Chloroniella'' c g * '' Corydalites'' c g * ''
Corydalus ''Corydalus'' is a genus of large flying insects in the Corydalidae family, commonly known as dobsonflies. They are endemic to North, Central and South America and there are about 35 known species. Members of the genus have wing lengths of up t ...
'' Latreille, 1802 i c g b – dobsonflies * '' Cratocorydalopsis'' c g * '' Cretochaulus'' c g * '' Ctenochauliodes'' c g * '' Dysmicohermes'' Munroe, 1953 i c g b * '' Eochauliodes'' c g * '' Jurochauliodes'' c g * '' Lithocorydalus'' c g * '' Madachauliodes'' c g * '' Neochauliodes'' c g * ''
Neohermes ''Neohermes'' is a genus of fishflies in the family Corydalidae The family Corydalidae contains the Megaloptera, megalopterous insects known as dobsonflies and fishflies. Making up about one dozen genera,See references in Haaramo (2008) they o ...
'' Banks, 1908 i c g b * '' Neoneuromus'' c g * '' Neurhermes'' c g * '' Neuromus'' g * '' Nevromus'' c g * ''
Nigronia Nigronia is a genus of megalopterous insects (dark fishflies) in the family of Corydalidae The family Corydalidae contains the Megaloptera, megalopterous insects known as dobsonflies and fishflies. Making up about one dozen genera,See referen ...
'' Banks, 1908 i c g b – dark fishflies * '' Nothochauliodes'' c g * ''
Orohermes ''Orohermes'' is a genus of fishflies in the family Corydalidae The family Corydalidae contains the Megaloptera, megalopterous insects known as dobsonflies and fishflies. Making up about one dozen genera,See references in Haaramo (2008) they o ...
'' Evans, 1984 i c g b * '' Parachauliodes'' c g * '' Platychauliodes'' c g * '' Platyneuromus'' Weele, 1909 i c g * ''
Protochauliodes ''Protochauliodes'' is a genus of fishflies in the family Corydalidae The family Corydalidae contains the Megaloptera, megalopterous insects known as dobsonflies and fishflies. Making up about one dozen genera,See references in Haaramo (2008) ...
'' Weele, 1909 i c g b * '' Protohermes'' van-der Weele, 1907 g * ''
Puri Puri () is a coastal city and a Nagar Palika, municipality in the state of Odisha in eastern India. It is the district headquarters of Puri district and is situated on the Bay of Bengal, south of the state capital of Bhubaneswar. It is als ...
'' c g * '' Sinochauliodes'' c g * '' Taeniochauliodes'' c g Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net


Footnotes


References


External links


Informative Corydalidae video
* * {{Taxonbar, from=Q1936061 Megaloptera Insect families