Cornus Stolonifera
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''Cornus sericea'', the red osier or red-osier dogwood, is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Cornaceae The Cornaceae are a cosmopolitan family of flowering plants in the order Cornales. The family contains approximately 85 species in two genera, ''Alangium'' and ''Cornus''. They are mostly trees and shrubs, which may be deciduous or evergreen, alt ...
, native to much of North America. It has sometimes been considered a synonym of the Asian species ''
Cornus alba ''Cornus alba'', the red-barked, white or Siberian dogwood, is a species of flowering plant in the family Cornaceae, native to Siberia, northern China and Korea. It is a large deciduous surculose (suckering) shrub that can be grown as a small ...
''. Other names include red brush, red willow,
USDA The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of com ...
NRCSbr>Plant Guide: REDOSIER DOGWOOD. May, 2006
/ref>Hilger, Inez (1951, repr. 1992) ''Chippewa Child Life and Its Cultural Background'', page 63Hart, Jeff, and Jacqueline Moore (1992). ''Montana—native plants and early peoples''
pages 38–39
Montana Historical Society.
redstem dogwood, redtwig dogwood, red-rood, American dogwood, creek dogwood, and western dogwood.


Description

In the wild, it most commonly grows in areas of rich, poorly drained soils, such as riparian zones and
wetlands A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...
, or in upland areas which receive more than 20 inches of precipitation annually. More uncommonly, it may be found in drier zones albeit at lesser abundance. Red osier dogwood is tolerant of flooding and has been known to survive up to seven years of water above root crown level. It occurs from sea level to 10,000 feet (3,000m), but in many areas is most common above 1,500 feet. It is a medium to tall
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
, growing 1.5–4 m tall and 3–5 m wide, spreading readily by underground
stolon In biology, stolons (from Latin '' stolō'', genitive ''stolōnis'' – "branch"), also known as runners, are horizontal connections between organisms. They may be part of the organism, or of its skeleton; typically, animal stolons are external s ...
s to form dense thickets. The branches and twigs are dark red, although wild plants may lack this coloration in shaded areas. The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are opposite, 5–12 cm long and 2.5–6 cm broad, with an ovate to oblong shape and an entire margin; they are dark green above and
glaucous ''Glaucous'' (, ) is used to describe the pale grey or bluish-green appearance of the surfaces of some plants, as well as in the names of birds, such as the glaucous gull (''Larus hyperboreus''), glaucous-winged gull (''Larus glaucescens''), g ...
below; fall color is commonly bright red to purple. Like all dogwoods, they have characteristic stringy white piths within the leaf stalks, which can be used for identification. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s are small (5–10 mm diameter), flat, umbella-like and dull white, in clusters 3–6 cm diameter. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
is a
globose A sphere () is a geometrical object that is a three-dimensional analogue to a two-dimensional circle. A sphere is the set of points that are all at the same distance from a given point in three-dimensional space.. That given point is the ce ...
white
berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
5–9 mm diameter. The Latin
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''sericea'' means "silky", referring to the texture of the leaves.


Ecology

Red osier dogwood provides food and cover for many species of mammals and birds. The stems and especially new shoots are browsed by moose, elk, bighorn sheep, mountain goats, beavers, and rabbits, while the fruits are an important autumn food source for bears, small mammals, and 47 different bird species. In winter, red osier dogwood is heavily browsed by
ungulates Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, cam ...
; in some areas use exceeds availability and individuals which have not been browsed are rare. The shrub is also important for nesting habitat and cover for a great variety of animals. ''Cornus sericea'' is shade tolerant but prefers intermediate to high light levels. It tolerates disturbance well, and appears early in both primary and secondary succession throughout its native range, but especially in floodplains and riparian zones. It thrives in fire-disturbed sites, sprouting from seeds or damaged shrubs. Although its conservation status is overall secure, it is considered vulnerable in
Iowa Iowa () is a state in the Midwestern region of the United States, bordered by the Mississippi River to the east and the Missouri River and Big Sioux River to the west. It is bordered by six states: Wisconsin to the northeast, Illinois to the ...
and critically imperiled in
Kentucky Kentucky ( , ), officially the Commonwealth of Kentucky, is a state in the Southeastern region of the United States and one of the states of the Upper South. It borders Illinois, Indiana, and Ohio to the north; West Virginia and Virginia to ...
and
Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...
.


Cultivation

''Cornus sericea'' is a popular ornamental shrub that is often planted for the red coloring of its twigs in the dormant season. The
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...
s 'Bud's Yellow', 'Flaviramea' with lime green stems, and 'Hedgerows Gold' (variegated foliage) have gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nort ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
(confirmed 2017). Like most dogwood species native to North America, ''C. sericea'' can be parasitized by the
dogwood sawfly ''Cornus'' is a genus of about 30–60 species of woody plants in the family Cornaceae, commonly known as dogwoods, which can generally be distinguished by their blossoms, berries, and distinctive bark. Most are deciduous trees or shru ...
, possibly leaving much of the plant devoid of leaves. A variety of pesticides are effective; however, hand-picking the larvae is also an option.


Uses

''Cornus sericea'' is frequently used for waterway bank erosion protection and restoration in the United States and Canada. Its root system provides excellent soil retention, it is hardy and provides an attractive shrub even when bare in winter, and its ability to be reproduced by cuttings makes it a low-cost solution for large-scale plantings. Some Plateau Indigenous tribes ate the berries to treat
colds The common cold or the cold is a virus, viral infectious disease of the upper respiratory tract that primarily affects the Respiratory epithelium, respiratory mucosa of the human nose, nose, throat, Paranasal sinuses, sinuses, and larynx. Si ...
and to slow bleeding.Moerman, Daniel E. (1998) "''Cornus sericea ssp. occidentallis''" ''Native American ethnobotany'' Timber Press, Portland, Oregon
page 178
Known as ''cansasa'' in
Lakota Lakota may refer to: *Lakota people, a confederation of seven related Native American tribes *Lakota language, the language of the Lakota peoples Place names In the United States: *Lakota, Iowa *Lakota, North Dakota, seat of Nelson County *Lakota ...
, the inner bark was also used by the Lakota and other Native Americans as "traditional tobacco", either by itself or in a mixture with other plant materials. Among the
Algonquian peoples The Algonquian are one of the most populous and widespread North American native language groups. Historically, the peoples were prominent along the Atlantic Coast and into the interior along the Saint Lawrence River and around the Great Lakes. T ...
such as the
Ojibwe The Ojibwe, Ojibwa, Chippewa, or Saulteaux are an Anishinaabe people in what is currently southern Canada, the northern Midwestern United States, and Northern Plains. According to the U.S. census, in the United States Ojibwe people are one of ...
, the smoking mixtures, known as
kinnikinnick Kinnikinnick is a Native American and First Nations herbal smoking mixture, made from a traditional combination of leaves or barks. Recipes for the mixture vary, as do the uses, from social, to spiritual to medicinal. Etymology The term "kinniki ...
, blended the inner bark with
tobacco Tobacco is the common name of several plants in the genus '' Nicotiana'' of the family Solanaceae, and the general term for any product prepared from the cured leaves of these plants. More than 70 species of tobacco are known, but the ...
, while more western tribes added it to the
bearberry Bearberries (indigenous kinnickinnick) are three species of dwarf shrubs in the genus ''Arctostaphylos''. Unlike the other species of ''Arctostaphylos'' (see manzanita), they are adapted to Arctic and Subarctic climates, and have a circumpolar d ...
leaf to improve the taste. The Ojibwe used red-osier dogwood bark as a dye by taking the inner bark and mixing it with other plants or minerals. The withies, or osiers, are used in basketry.


Subspecies

It is a variable species, with two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
commonly accepted: *''Cornus sericea'' subsp. ''sericea'' – throughout the range of the species. Shoots and leaves hairless or finely pubescent; flower petals 2–3 mm. *''Cornus sericea'' subsp. ''occidentalis'' (Torr. & A.Gray) Fosberg – western North America. Shoots and leaves densely pubescent; flower petals 3–4.5 mm.


Distribution

It is
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
throughout boreal and temperate zones in northern and western North America from Alaska east to Newfoundland, south to Durango and Nuevo León in the west, and Illinois and Virginia in the east. *''Cornus sericea'' L. has been recorded from counties Antrim and Londonderry in
Northern Ireland Northern Ireland ( ga, Tuaisceart Éireann ; sco, label= Ulster-Scots, Norlin Airlann) is a part of the United Kingdom, situated in the north-east of the island of Ireland, that is variously described as a country, province or region. Nort ...
.Hackney, P. 1992. "Stewart & Corry's Flora of the North-east of Ireland." Third edition. Institute of Irish Studies, The Queen's University of Belfast. (HB)


Gallery

Image:Cornus sericea habit.jpg, shrub Image:Cornus sericea fall.jpg, shrub, fall Image:Cornus sericea winter.jpg, shrub, winter Image:Cornus Stolonifera garden show 1.jpg, twigs Image:Cornus sericea leaf.jpg, leaf Image:Cornus sericea twig.jpg, twig and leaf Image:Cornus sericea leaves and inflorescence 2003-08-11.jpg, flowers Image:Redtwigdog.jpg, flowers Image:Cornus sericea flower.jpg, flowers Image:Cornus sericea HdB.png, ''Cornus: specimen botanicum sistens descriptiones et icones specierum corni minus cognitarum'' Image:Cornus sericea seed.png, seed


References

*


External links

*
Jepson Flora of California: ''Cornus sericea''subsp. ''sericea''subsp. ''occidentalis''


* ttp://plants.usda.gov/java/profile?symbol=COSE16 NRCS: USDA Plants Profile: ''Cornus sericea''* {{Authority control sericea Plants used in traditional Native American medicine Flora of Subarctic America Flora of Eastern Canada Flora of Western Canada Flora of the Northwestern United States Flora of the North-Central United States Flora of the Northeastern United States Flora of the Southwestern United States Flora of the South-Central United States Flora of the Southeastern United States Flora of Northeastern Mexico