Coralloidoolithus
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''Coralloidoolithus'' is an oogenus of dinosaur egg from the Tiantai Basin in
Zhejiang Province Zhejiang ( or , ; , also romanized as Chekiang) is an eastern, coastal province of the People's Republic of China. Its capital and largest city is Hangzhou, and other notable cities include Ningbo and Wenzhou. Zhejiang is bordered by Jiangs ...
, containing a single known oospecies ''C. shizuiwanensis''. Formerly, it was classified in the oogenus '' Paraspheroolithus''; however, it was considered sufficiently different to be classified in its own genus. ''C. shizuiwanensis'' is similar to ''
Stalicoolithus ''Stalicoolithus'' is an oogenus of dinosaur egg from the Sanshui and Chichengshan Formations of the Tiantai Basin in Zhejiang Province. It is known from a single, complete fossil egg, notable for its spherical shape.Wang Q, Wang X L, Zhao Z K, ...
'', leading to their classification in the same family,
Stalicoolithidae Stalicoolithidae is an oofamily of fossil eggs.Wang Q, Wang X L, Zhao Z K, and Jiang Y G. (2012).A new oofamily of dinosaur egg from the Upper Cretaceous of Tiantai Basin, Zhejiang Province, and its mechanism of eggshell formation Chinese Scienc ...
.Wang Q, Wang X L, Zhao Z K, and Jiang Y G. (2012).
A new oofamily of dinosaur egg from the Upper Cretaceous of Tiantai Basin, Zhejiang Province, and its mechanism of eggshell formation
Chinese Science Bulletin ''Science Bulletin'' () is a multidisciplinary scientific journal co-sponsored by the Chinese Academy of Sciences and the National Natural Science Foundation of China. It is published by Elsevier on behalf of Science in China Press and focuses on ...
. 57: 3740-3747. doi: 10.1007/s11434-012-5353-2


History

Xixia County was first excavated for fossils by a Geological Team of the Henan Geological Bureau in 1974. Fossils continued to be excavated there, but were not extensively studied until 1994 when Fang ''et al.'' received funding from the
National Natural Science Foundation of China The National Natural Science Foundation of China (NSFC; ) is an organization directly affiliated to China's State Council for the management of the National Natural Science Fund. History NSFC was founded in February 1986 by theoretical chemist Tan ...
. Under this sponsorship, with support of the Geological Profession Foundation and the Geological Science Item Foundation of MGMR, a multidisciplinary team of Chinese paleontologists took on the task of analyzing the geology and paleontology of these eggs. In 1998, Fang ''et al.'' described numerous new ootaxa, including the oospecies ''" Paraspheroolithus" shizuiwanensis''. Further specimens were uncovered in
Tiantai Tiantai or T'ien-t'ai () is an East Asian Buddhist school of Mahāyāna Buddhism that developed in 6th-century China. The school emphasizes the ''Lotus Sutra's'' doctrine of the "One Vehicle" (''Ekayāna'') as well as Mādhyamaka philosophy, ...
in 2000; these were referred to ''"P."'' cf. ''shizuiensis''. In 2012 multiple complete and near-complete eggs were discovered in Tiantai by the Chinese paleontologists Wang Qiang, Wang Xiaolin, Zhao Zikui, and Jiang Yan'gen. They named a new oogenus, ''Coralloidoolithus,'' for ''P. shizuiwanensis'', and referred several of their newly discovered specimens to ''C. shizuiwanensis''. They also synonymized this oospecies with the
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
n ''
Dendroolithus ''Dendroolithus'' is an oogenus of Dendroolithid dinosaur egg found in the late Cenomanian Chichengshan Formation ( Tiantai Group), in the Gong-An-Zhai and Santonian Majiacun Formations of China and the Maastrichtian Nemegt and Campanian Bar ...
microporosus'' and ''Spheroolithus'' oosp. from
South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korea, Korean Peninsula and sharing a Korean Demilitarized Zone, land border with North Korea. Its western border is formed ...
based on similarities in microstructure.


Description

The first specimens discovered were only fragments, but based on impressions they were estimated to be in diameter. The complete eggs discovered in Tiantai are larger, ranging from in length by wide. They are roughly spherical. Wang ''et al.'' (2012) described them as having a rough surface, but the original ''"P." shizuiwanensis'' specimens were noted by Fang ''et al.'' to have a smooth surface. ''Dendroolithus macroporosus'' and the ''Spheroolithus'' specimens referred to ''C. shizuiwanensis'' are also smooth-shelled. ''Coralloidoolithus'' shell is typically about 2.6 mm thick. It has similar microstructures to the other stalicoolithids; it has a very thin cone layer (the inner layer of the eggshell), making up only about 1/10 of the eggshell's thickness, and its columnar layer is divided into three zones. Secondary shell units make up the outermost of these zones and also fill many of the pore canals. The middle zone is characterized by light and dark bands and the innermost zone of the columnar layer has horizontal growth lines. Its pores are irregular and meandering. ''Coralloidoolithus'' can be distinguished from ''Stalicoolithus'' by having a thinner eggshell and by having fewer secondary shell units. '' Shixingoolithus'' is larger than ''Coralloidoolithus'' and has a thicker cone layer.


Distribution

''Coralloidoolithus shizuiwanensis'' is found at several
Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
sites, including at the Sangping Formation in Xixia County,
Henan Henan (; or ; ; alternatively Honan) is a landlocked province of China, in the central part of the country. Henan is often referred to as Zhongyuan or Zhongzhou (), which literally means "central plain" or "midland", although the name is al ...
and at the Chichengshan Formation Tiantai County,
Zhejiang Zhejiang ( or , ; , also romanized as Chekiang) is an eastern, coastal province of the People's Republic of China. Its capital and largest city is Hangzhou, and other notable cities include Ningbo and Wenzhou. Zhejiang is bordered by Jiang ...
. Also, specimens found in the
Gobi Desert The Gobi Desert (Chinese: 戈壁 (沙漠), Mongolian: Говь (ᠭᠣᠪᠢ)) () is a large desert or brushland region in East Asia, and is the sixth largest desert in the world. Geography The Gobi measures from southwest to northeast an ...
and in Bosung County, Chullanam-do Province,
South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korea, Korean Peninsula and sharing a Korean Demilitarized Zone, land border with North Korea. Its western border is formed ...
have been referred to this oospecies.


Classification

''Coralloidoolithus'' is a
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
oogenus containing only ''C. shizuiwanensis''. It is classified alongside ''Stalicoolithus'' and ''Shixingoolithus'' in the oofamily Stalicoolithidae.


See also

*
List of dinosaur oogenera Egg fossils are the fossilized remains of eggs laid by ancient animals. As evidence of the physiological processes of an animal, egg fossils are considered a type of trace fossil. Under rare circumstances a fossil egg may preserve the remains of t ...


References

{{Taxonbar, from=Q22286541 Stalicoolithids Dinosaur reproduction Cretaceous China Fossils of China Fossil parataxa described in 2012