The copper underwing, humped green fruitworm or pyramidal green fruitworm (''Amphipyra pyramidea'') is a

moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...

of the family Noctuidae. The species was first described by

Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his Nobility#Ennoblement, ennoblement in 1761 as Carl von Linné#Blunt, Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalise ...

in his 1758 10th edition of ''Systema Naturae''.

Distribution

This species can be found across the

Palaearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...

region including Europe,

North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...

, the Near East,

Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...

, southern

Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive region, geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a ...

, northern

India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...

Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic o ...

and Japan. It is rather common over the southern half of Britain.

Description

This species has a

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan o ...

of 47–54 mm, the female usually slightly larger than the male. The forewings are brown marked with paler fascia and a pale, dark-centred stigma. The hindwings are a rich bright copper colour (hence the common name of "copper underwing"). It is very similar to Svensson's copper underwing (''Amphipyra berbera'') but identification is usually fairly straightforward by looking at some markings on the forewings, the detail of the labial palps and at the underside of the hindwings. This species has a pale area in the centre of the hindwings, contrasting with much darker marginal areas, while in ''A. berbera'' the whole underwing is more or less uniform in colour. See Townsend et al. The

larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...

is green with white markings and a pointed hump at the rear end.

Differentiation of ''A. pyramidea'' from ''A. berbera''

File:Amphipyra pyramidea - differenciation from Amphipyra berbera 01.jpg, ''A. pyramidea'' - red arrow points out a difference File:Amphipyra pyramidea - differenciation from Amphipyra berbera 02.jpg, ''A. berbera'' - red arrows point out a difference File:Amphipyra pyramidea - berbera larvae.jpg, Caterpillars of ''A. pyramidea'' (top) and ''A. berbera''

Biology

It is a

univoltine Voltinism is a term used in biology to indicate the number of broods or generations of an organism in a year. The term is most often applied to insects, and is particularly in use in sericulture, where silkworm varieties vary in their voltinism. ...

species. Hatching season is as early as June in some climates but they usually hatch July to October. Eggs are laid in deciduous trees. Larva can be found as early as April in some climates but usually emerge in May–June. ''A pyramidea'' flies at night from August to October and is attracted to light and strongly to sugar. It feeds on a variety of

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

s and shrubs, including ash tree ('' Fraxinus''), privet (''

Ligustrum A privet is a flowering plant in the genus ''Ligustrum''. The genus contains about 50 species of erect, deciduous or evergreen shrubs, sometimes forming small or medium-sized trees, native to Europe, north Africa, Asia, many introduced and nat ...

''), honeysuckle (''

Lonicera Honeysuckles are arching shrubs or twining vines in the genus ''Lonicera'' () of the family Caprifoliaceae, native to northern latitudes in North America and Eurasia. Approximately 180 species of honeysuckle have been identified in both contin ...

''), apple (''

Malus ''Malus'' ( or ) is a genus of about 30–55 species of small deciduous trees or shrubs in the family Rosaceae, including the domesticated orchard apple, crab apples, wild apples, and rainberries. The genus is native to the temperate zone ...

''), oak (''

Quercus An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ...

''), '' Rhododendron'',

rose A rose is either a woody perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred species and tens of thousands of cultivars. They form a group of plants that can be ...

, wild service tree (''

Sorbus ''Sorbus'' is a genus of over 100 species of trees and shrubs in the rose family, Rosaceae. Species of ''Sorbus'' (''s.l.'') are commonly known as whitebeam, rowan ( mountain-ash) and service tree. The exact number of species is disputed depe ...

'') and lilac (''

Syringa ''Syringa'' is a genus of 12 currently recognized species of flowering woody plants in the olive family or Oleaceae called lilacs. These lilacs are native to woodland and scrub from southeastern Europe to eastern Asia, and widely and commonly ...

'').Mazzei, Paolo; Morel, Daniel & Panfili, Raniero
''Moths and Butterflies of Europe and North Africa''
/ref> The species overwinters as an

egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...

. # ''The flight season refers to the

British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...

. This may vary in other parts of the range.''

Gallery

File: Amphipyra pyramidea larva.jpg, Larva File: Noctuidae - Amphipyra pyramidea.JPG, Moth File:Amphipyra pyramidea.jpg, Mounted specimen File: F Nemos OBA Amphipyra pyramidea.jpg, Illustration

References

* *

External links

''Butterflies & Moths of Switzerland''''Lepiforum e. V.''
Includes photo of genitalia.

{{Taxonbar, from=Q735875 Amphipyrinae Moths of North America Moths of Europe Moths of Asia Moths of Africa Moths described in 1758 Taxa named by Carl Linnaeus