Conotrachelus Posticatus
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The ''Conotrachelus posticatus'' is a species of
true weevil True most commonly refers to truth, the state of being in congruence with fact or reality. True may also refer to: Places * True, West Virginia, an unincorporated community in the United States * True, Wisconsin, a town in the United States * T ...
within the
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
family
Curculionidae The Curculionidae are a family of weevils, commonly called snout beetles or true weevils. They are one of the largest animal families, with 6,800 genera and 83,000 species described worldwide. They are the sister group to the family Brentidae. T ...
. ''C''. ''posticatus'' is found in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. It is on average long, and it is present in North America (particularly in
Minnesota Minnesota () is a state in the upper midwestern region of the United States. It is the 12th largest U.S. state in area and the 22nd most populous, with over 5.75 million residents. Minnesota is home to western prairies, now given over to ...
,
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
, and
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
) all the way to
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
. This species of ''Conotrachelus'' breeds in
acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera ''Quercus'' and '' Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and borne ...
s; as a result, its
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e are found within acorns and their larval diet includes acorns. Their growing presence within
oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...
trees has been studied and their effect on oak trees shows that they attack these trees by destroying the seeds within the tree. At times, the seeds can be found to be partially damaged, and requires further studying to understand the activity of the species in relation to oak trees.


Background description

''Conotrachelus posticatus'' are more prevalent throughout the regions of North America, especially in areas such as Minnesota to Texas and Panama. It has a long nose that bears some resemblance to the trunk of an elephant. The nose has a chewing mouth piece at its end, allowing for it to process its food. ''C. posticatus'' is brown and/or black in color. ''C. posticatus'' use acorns as a food source in both its adult and larval stages. Acorns can also be used as nurseries for these particular beetles. Females tend to lay their eggs into the seed of oak trees (in the tissues of green acorns) and larvae feed on the inside of the seed. ''C. posticatus'' breeds in acorns of nine different species of oak trees, and the eggs are often deposited in damaged acorns. Their larval stage lasts between 10-30 days, and the second winter of ''C. posticatus'' tends be spent under leaves on the ground as an adult. ''C. posticatus'' seems to be related to that of the '' C. naso'' beetle in range and in how the eggs are deposited. It remains hidden from its predators during the wintertime inside of soil, after it has chewed into the hole of an acorn seed. The natural relationship between the ''C. posticatus'' and the acorn seed can pose a threat to the oak trees. Predators of ''C. posticatus'' reduce the threat it poses to oak trees. One feature of ''C. posticatus'' is that it uses a stridulatory mechanism to produce sound. This is done by scraping the elytral stridulitra and the plectra together to produce frequencies. There are demonstrable differences that exist between the stridulations of this beetle and other beetles within its genus. Individual beetles are capable of several different speeds and frequencies of stridulations. Sound production can be induced by agitation or rough handling of the beetles.


Life history

The article, "Biology and Life History of Acorn-Infesting Weevils of the Genus Conotrachelus (Coleoptera: Curculionidae)" by Lester P. Gibson, analyzes the life history of the ''C. naso'' LeConte beetle, the ''C. posticatus'' Boheman beetle, and the '' C. carinifer'' Casey beetle. This study had begun in October 1960 with
Ohio Ohio () is a state in the Midwestern region of the United States. Of the fifty U.S. states, it is the 34th-largest by area, and with a population of nearly 11.8 million, is the seventh-most populous and tenth-most densely populated. The sta ...
acorns, and unfolded two years later in 1962 to include more oak trees in the U.S. When looking particularly at the life history of ''C. posticatus'', it was revealed that the larvae of the ''C. posticatus'' species had gone through 5
instar An instar (, from the Latin '' īnstar'', "form", "likeness") is a developmental stage of arthropods, such as insects, between each moult (''ecdysis''), until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow or ass ...
s, where they emerged in 14 days from
bur oak ''Quercus macrocarpa'', the bur oak or burr oak, is a species of oak tree native to eastern North America. It is in the white oak section, ''Quercus'' sect. ''Quercus'', and is also called mossycup oak, mossycup white oak, blue oak, or scrub oa ...
acorns and 30 days from the black acorns that were conducive to this study. It was also noted that the "emergence method, cell formation, and prepupal and pupal activity patterns are similar to those of ''C. naso'' larvae." Additionally, the study found that ''C. posticatus'' larvae overwinter within the soil, but the adults tend to hibernate underneath leaves. This showed the researchers that the adult ''C. posticatus'' emerges in the late spring and early summer, where it " oviposits in acorns, and passes the second winter as an adult." Nevertheless, there were certain limitations to what the researchers were able to find out about the species in terms of how long the adult can live after surviving its second winter. The researchers also noted that there was very little information given on the biology of the ''C. posticatus'', but that the species was similar to that of the ''C. naso'' species as they both are found in areas like Panama. Much of this study was done to examine the
longevity The word " longevity" is sometimes used as a synonym for "life expectancy" in demography. However, the term ''longevity'' is sometimes meant to refer only to especially long-lived members of a population, whereas ''life expectancy'' is always d ...
and adult emergence of ''C. posticatus'', and their breeding activity in
Crataegus ''Crataegus'' (), commonly called hawthorn, quickthorn, thornapple, Voss, E. G. 1985. ''Michigan Flora: A guide to the identification and occurrence of the native and naturalized seed-plants of the state. Part II: Dicots (Saururaceae–Cornacea ...
fruits, fresh
hickory Hickory is a common name for trees composing the genus ''Carya'', which includes around 18 species. Five or six species are native to China, Indochina, and India (Assam), as many as twelve are native to the United States, four are found in Mexi ...
nuts, and black oak acorn meats. The results showed that most of the ''C. posticatus'' beetles would feed, lay their eggs, and produce the most larvae in the black oak acorn meats. Overall, this species seems to adapt, utilize, and occupy acorns as a resource for survival.


The ''Conotrachelus'' and acorns

The species of ''Conotrachelus'' in all stages of its life (larvae, pupae, and adulthood), breeds within acorns specifically in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
and
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
. The physical description and behavior of the ''Conotrachelus'' species is inspected in the research study, "Systematics of the Acorn-Infesting Weevils Conotrachelus naso, C. carinifer, and C. posticatus (Coleoptera: Curculionidae)", by Lester P. Gibson. This research study examines a detailed view of the larvae, pupae, and the adulthood of the ''C. naso'', ''C. carinifer'', and ''C. posticatus''. In particular, the ''C. posticatus'', in its adult stage is described as its color being "dark reddish-brown", its head being "densely
punctate Punctum, plural puncta, adjective punctate, is an anatomical term for a sharp point or tip. It may also refer to: Medical *Lacrimal punctum, a minute opening on the margins of the eyelids that collect tears produced by the lacrimal glands *Blin ...
", and its special characteristics, including its dense
prothorax The prothorax is the foremost of the three segments in the thorax of an insect, and bears the first pair of legs. Its principal sclerites (exoskeletal plates) are the pronotum (dorsal), the prosternum (ventral), and the propleuron (lateral) on ea ...
, its legs, and its genitalia. Much of this physical description of the adult weevil beetle is to explain how and why this beetle is able to take hold of and utilize acorns to its own advantage. For instance, in the article, "Acorn Weevil (Coleoptera: Curculionidae) Predation Dynamics in a
Mississippi Mississippi () is a state in the Southeastern region of the United States, bordered to the north by Tennessee; to the east by Alabama; to the south by the Gulf of Mexico; to the southwest by Louisiana; and to the northwest by Arkansas. Miss ...
Bottomland
Hardwood Hardwood is wood from dicot trees. These are usually found in broad-leaved temperate and tropical forests. In temperate and boreal latitudes they are mostly deciduous, but in tropics and subtropics mostly evergreen. Hardwood (which comes from ...
Forest", by Justin P. Williams and Tracy S. Hawkins, illustrates how there is a sense of
predation Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
in acorns for individual trees that produce such acorns, because of the weevil beetle. The researchers had placed cone emergence traps in a periodically flooded forest to obtain weevil population
parameter A parameter (), generally, is any characteristic that can help in defining or classifying a particular system (meaning an event, project, object, situation, etc.). That is, a parameter is an element of a system that is useful, or critical, when ...
estimate Estimation (or estimating) is the process of finding an estimate or approximation, which is a value that is usable for some purpose even if input data may be incomplete, uncertain, or unstable. The value is nonetheless usable because it is der ...
s and understanding tree-to-tree variability in acorn predation rates. By having done these parameters to analyze the predation rates of weevils, it can help to combat the reduction of food resources that wildlife needs, and preventing these beetles from replenishing seedling placement. The results revealed that the ''Conotrachelus'' weevils were seen as "secondary acorn predators", yet they were also seen as helping to regenerate bottomland forests from being utterly gone, because they are able to "outwinter as adults and infest 20-65% of spring-germinating acorns on the soil surface". Even though the ''Conotrachelus'' beetle may cause harm to acorns, it can also bring about more abundance in the quantity of the forest.


Global effects of the ''C. posticatus'' beetle

The ''C. posticatus'' beetle has been regarded as one of the seven species of ''Curculionidae'' to cause a threat to multimillion-dollar
avocado The avocado (''Persea americana'') is a medium-sized, evergreen tree in the laurel family (Lauraceae). It is native to the Americas and was first domesticated by Mesoamerican tribes more than 5,000 years ago. Then as now it was prized for i ...
industries. Within the research study, "On the identity of a U.S. intercepted Conotrachelus Dejean (Coleoptera: Curculionidae) with avocado (Persea americana)", by Maria Lourdes Chamorro and Maxwell V. L. Barclay, it strives to analyze the identification of non-native weevils and their effect on local agriculture. Firstly, this study begins by comparing adult weevils with "male and female type specimens of ''C. lobatus'' Champion, ''C. squamifrons'' Champion and ''C. scoparius'' Champion." to be able to help those that are growers on agricultural lands to absorb and understand all of the organisms' biology for better preventative measures against the growing threats from the species of weevil beetles. Further, this study found that the ''C. posticatus'' most closely looks similar to that of the ''Conotrachelus lobatus'', (where this particular species is found North America, north of
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
). Other notable physical descriptions regarding the ''C. posticatus'' is that, this species, amongst others related species (''C. carinifer'' Casey and ''C. naso'' LeConte) "all have…longitudinal median prothoracic carina;
mesosternum The mesothorax is the middle of the three segments of the Thorax (arthropod anatomy), thorax of Hexapoda, hexapods, and bears the second pair of legs. Its principal sclerites (exoskeletal plates) are the mesonotum (Dorsum (biology), dorsal), the m ...
with anterolateral angles truncate and prominent". This similarity amongst these species can further prove to show how common it may be to mix up the different beetles and categorize them as the same beetle due to their physiology. When looking at the discussion section of this study, the identification of the ''C. lobatus'' is a non-native species with avocados and can frequently be mistaken for the U.S. native ''C. posticatus''. Due to this misconception between both of the species, it can introduce non-native species into agricultural systems that may not be able to thrive under such conditions. The results had concluded that, ''posticatus'' individuals from
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
differ from those in the U.S. in having ventral segments less closely punctured." The nature of this physical relationship between the differing species of weevil beetles illustrates a greater importance of understanding how invasive species can cause for harm to an environment, and ecosystem, and even at times to human health.


References


Further reading

* * Molytinae Articles created by Qbugbot Beetles described in 1837 {{Molytinae-stub