Condorraptor Scale
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''Condorraptor'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
megalosauroid Megalosauroidea (meaning 'great/big lizard forms') is a superfamily (or clade) of tetanuran theropod dinosaurs that lived from the Middle Jurassic to the Late Cretaceous period. The group is defined as ''Megalosaurus bucklandii'' and all taxa sha ...
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
. Its genus name means 'robber from Cerro Condor', referencing a nearby village, while its species name, ''currumili'', is named after Hipolito Currumil, the landowner and discoverer of the locality. It was among the earliest large South American theropods, having been found in
Lower Jurassic The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-Jurassic extinction event, 201.3 Ma  ...
strata of the
Cañadón Asfalto Formation The Cañadón Asfalto Formation is a Lower Jurassic to Late Jurassic geologic formation, from the Jurassic period of the Mesozoic Era. Its age is controversial, uranium-lead dating of the volcanic tuff beds having given various different ages. A ...
in the
Cañadón Asfalto Basin The Cañadón Asfalto Basin ( es, Cuenca de Cañadón Asfalto) is an irregularly shaped sedimentary basin located in north-central Patagonia, Argentina. The basin stretches from and partly covers the North Patagonian Massif in the north, a high for ...
of
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
, described in 2005, is ''Condorraptor currumili''. It is based on a
tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
, with an associated partial skeleton that may belong to the same individual. Initially described as a basal tetanuran, Benson (2010) found it to be a piatnitzkysaurid megalosauroid and the sister taxon of ''
Piatnitzkysaurus ''Piatnitzkysaurus'' ( meaning "Piatnitzky's lizard") is a genus of megalosauroid theropod dinosaur that lived approximately 179 to 177 million years ago during the lower part of the Jurassic Period in what is now Argentina. ''Piatnitzkysaurus'' ...
,'' a finding supported by later studies.


Description

The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
of ''Condorraptor'' is MPEF-PV 1672, a left tibia. Additional remains (MPEF-PV 1673 through 1697 and MPEF-PV 1700 through 1705) have also been referred to the species, including
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
e, teeth, rib and chevron fragments, partial hip bones,
femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with ...
s, a
metatarsal The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the med ...
IV, and a pedal phalanx. All of these remains were from the same locality of the holotype and likely represent the same individual. In 2007, various media outlets reported that an articulated skeleton of this species was discovered by a team led by Oliver Rauhut, but this find has not been described or referenced in literature. Also in 2007, Rauhut described a fragmentary partial skull, MPEF 1717, from the Canadon Asfalto Formation. Due to the skull's size, locality,
tetanuran Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, tyrannosauroids, ornithomimosaurs, compsognathids and maniraptorans (including birds). Tetanurans ar ...
characteristics, and differences from the cranial material of ''Piatnitzkysaurus'', it is possible that it belongs to ''Condorraptor''. The type specimen was a juvenile that was about 4.5 metres long and it weighed about 200 kg. In 2016 Molina-Pérez and Larramendi estimated the probable adult specimen (MPEF 1717) at 8.5 meters and 1.65 tonnes. ''Condorraptor'' is notably similar to another theropod from the same formation, ''Piatnitzkysaurus''. Unique among tetanurans, these two share a flat anterior surface of the anterior
presacral In human anatomy, the presacral space is inside the pelvis, behind the rectum and in front of the coccyx and sacrum. Normally it is empty, or it contains a pocket of fat. It is usually covered by sigmoid colon. Clinical significance The presacral ...
centra. However, it can be distinguished from ''Piatnitzkysaurus'' and other megalosauroids by several diagnostic features. Although some features considered diagnostic by the original description were later shown to be present in other megalosauroids, several features are still only known in ''Condorraptor''. These include: * A pleurocoel in the anterior cervical vertebrae located immediately posterodorsal to the parapophysis. * A shallow depression on the lateral surface of the tibia at the base of the
cnemial crest The cnemial crest is a crestlike prominence located at the front side of the head of the tibiotarsus or tibia in the legs of many mammals and reptiles (including birds and other dinosaurs). The main extensor muscle of the thigh In human anatomy ...
. * A metatarsal IV with a distinct dorsal step between the shaft and the distal articular facet. In addition, ''Condorraptor'' differs from ''Piatnitzkysaurus'' by the shape of the underside of its sacral centra. In ''Condorraptor'', the second centra has a broad and flat base while the third is gently concave. In ''Piatnitzkysaurus'', the second centra's base is smoothly rounded while the third's is flat along its midline.


Classification

The most basal clade within
Megalosauroidea Megalosauroidea (meaning 'great/big lizard forms') is a superfamily (or clade) of tetanuran theropod dinosaurs that lived from the Middle Jurassic to the Late Cretaceous period. The group is defined as ''Megalosaurus bucklandii'' and all taxa sha ...
contains ''Condorraptor'', ''
Marshosaurus ''Marshosaurus'' is a genus of medium-sized carnivorous theropod dinosaur, belonging to the Megalosauroidea, from the Late Jurassic Morrison Formation of Utah and possibly Colorado. Description ''Marshosaurus'' was medium-sized for a theropod. ...
'', ''
Piatnitzkysaurus ''Piatnitzkysaurus'' ( meaning "Piatnitzky's lizard") is a genus of megalosauroid theropod dinosaur that lived approximately 179 to 177 million years ago during the lower part of the Jurassic Period in what is now Argentina. ''Piatnitzkysaurus'' ...
'' and ''
Xuanhanosaurus ''Xuanhanosaurus'' (meaning "Xuanhan lizard") is a genus of theropod dinosaur that lived during the Middle Jurassic of China, around 167.7 to 161.2 million years ago. Discovery The type species ''Xuanhanosaurus qilixiaensis'' was named by Don ...
''. The next most basal clade comprises ''
Chuandongocoelurus ''Chuandongocoelurus'' ( ) is a genus of carnivorous tetanuran theropod dinosaur from the Jurassic of China. Discovery and naming The type species ''Chuandongocoelurus primitivus'' was first described and named by Chinese paleontologist He ...
'' and ''
Monolophosaurus ''Monolophosaurus'' ( ; meaning "single-crested lizard") is an extinct genus of tetanuran theropod dinosaur from the Middle Jurassic Shishugou Formation in what is now Xinjiang, China.Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up ...
''. However, the affiliation of these clades with Megalosauroidea is poorly supported by tree support metrics, and it is possible that they will be classified outside of Megalosauroidea by future analyses.


References

{{Portal bar, Dinosaurs Megalosaurs Dinosaur genera Aalenian life Bajocian life Middle Jurassic dinosaurs of South America Jurassic Argentina Fossils of Argentina Cañadón Asfalto Formation Fossil taxa described in 2005